Tethysaurus
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Tethysaurus
''Tethysaurus'' is an extinct genus of tethysaurine mosasauroid from the Early Turonian (Late Cretaceous) period. The only species is ''Tethysaurus nopcsai''. Discovery The name means "Tethys' lizard of Nopcsa", a reference to the Greek goddess of the sea Tethys (also the name of the Tethys Ocean, an ancient sea between southern Europe and northern Africa) and to the Hungarian paleontologist Baron Ferenc Nopcsa, who made pioneering studies on Adriatic aquatic squamates. It was found in the Akrabou Formation, near the villages of Tadirhourst and Asfla in the region of Goulmima, Errachidia Province, in Morocco, with three referred specimens that included a nearly complete articulated skull, mandible, vertebrae and portions of the appendicular skeleton. The diagnosis after Bardet ''et al''. is "(...) prefrontal strongly vaulted in anterior view; parietal exhibits a triangular table ending posteriorly in two pointed pegs overlying the supraoccipital; jugal with a large and wide ...
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Tethysaurine
The Tethysaurinae are a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the subfamily are informally and collectively known as "tethysaurines" and have been recovered from North America and Africa. Only two tethysaurine genera are known, ''Pannoniasaurus'' and ''Tethysaurus''. The genera ''Yaguarasaurus'' and ''Russellosaurus'' were previously considered tethysaurines until they were grouped with '' Romeosaurus'' in the new subfamily Yaguarasaurinae. A possible member of this clade (subfamily) is a mosasaur specimen known from a maxilla fragment, found in 1960 in the Czech Republic (then Czechoslovakia), in Dolní Újezd near Litomyšl. Like the closely related yaguarasaurines, all tethysaurines were plesiopedal (meaning primitive and not as well adapted to marine life as later mosasaurs). They generally retained relatively small sizes compared to later giant mosasaurs. The tethysaurines appeared during the Turonian and went extinct in the S ...
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Tethysaurus Nopcsai 45
''Tethysaurus'' is an extinct genus of tethysaurine mosasauroid from the Early Turonian (Late Cretaceous) period. The only species is ''Tethysaurus nopcsai''. Discovery The name means "Tethys' lizard of Nopcsa", a reference to the Greek goddess of the sea Tethys (also the name of the Tethys Ocean, an ancient sea between southern Europe and northern Africa) and to the Hungarian paleontologist Baron Ferenc Nopcsa, who made pioneering studies on Adriatic aquatic squamates. It was found in the Akrabou Formation, near the villages of Tadirhourst and Asfla in the region of Goulmima, Errachidia Province, in Morocco, with three referred specimens that included a nearly complete articulated skull, mandible, vertebrae and portions of the appendicular skeleton. The diagnosis after Bardet ''et al''. is "(...) prefrontal strongly vaulted in anterior view; parietal exhibits a triangular table ending posteriorly in two pointed pegs overlying the supraoccipital; jugal with a large and wide ...
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Yaguarasaurus
''Yaguarasaurus'' is an extinct genus of mosasauroid from the Late Cretaceous (Turonian) period of Colombia, South America. The remains discovered (an articulated skull, some vertebrae and ribs) were defined as a new genus and species of mosasaurid, ''Yaguarasaurus columbianus'', by the Colombian paleontologist María Páramo, former director of the Museo de Geología José Royo y Gómez of INGEOMINAS in Bogotá. The first fossils remains of this animal suggested a cranial length of and a total length of ; an additional skull that measures long implies a larger size. This reptile is a member of the family of marine lizards Mosasauridae characteristic of Middle and Upper Cretaceous, with global distribution, but in South America known only through isolated remains (Price, 1957, Pierce and Welles, 1959 ; Bonaparte, 1978; Ameghino, 1918). This mosasaur discovered in Yaguará, was at the moment of discovery the most complete material known in South America. Etymology The remain ...
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Russellosaurus
''Russellosaurus'' is an extinct genus of tethysaurine mosasauroid from the Late Cretaceous of North America. The genus was described from a skull discovered in an exposure of the Arcadia Park Shale (lower Middle Turonian) at Cedar Hill, Dallas County in south-central Texas, United States. The skull (SMU 73056, Shuler Museum of Paleontology, Southern Methodist University) was found in 1992 by a member of the Dallas Paleontological Society, who then donated to the museum. Other fragmentary specimens of ''Russelosaurus'' have been recovered from the slightly older Kamp Ranch Limestone at two other localities in the Dallas area. Etymology The type species, ''R. coheni'', was named for the amateur fossil collector who discovered SMU 73056, and the genus name honours paleontologist Dale A. Russell for his extensive work on mosasaurs ("Russell's lizard"). This is the second species of mosasaur to have been named for Russell, the first being ''Selmasaurus russelli'' (Wright and Shannon, ...
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Mosasauroid
Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Ancient Greek, Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. They belong to the order Squamata, which includes lizards and snakes. Mosasaurs probably evolved from an extinct group of aquatic lizards known as Aigialosauridae, aigialosaurs in the Late Cretaceous, Earliest Late Cretaceous with 42 described genera. During the last 20 million years of the Cretaceous period (Turonian–Maastrichtian ages), with the extinction of the ichthyosaurs and Pliosauridae, pliosaurs, mosasaurs became the dominant marine predators. They themselves became extinct as a result of the Cretaceous–Paleogene extinction event, K-Pg event at the end of the Cretaceous period, about 66 million years ago. Description Mosasaurs breathed air, were powerful swimmers, and were well-adapted to livi ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Morocco
Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to the east, and the disputed territory of Western Sahara to the south. Mauritania lies to the south of Western Sahara. Morocco also claims the Spanish exclaves of Ceuta, Melilla and Peñón de Vélez de la Gomera, and several small Spanish-controlled islands off its coast. It spans an area of or , with a population of roughly 37 million. Its official and predominant religion is Islam, and the official languages are Arabic and Berber; the Moroccan dialect of Arabic and French are also widely spoken. Moroccan identity and culture is a mix of Arab, Berber, and European cultures. Its capital is Rabat, while its largest city is Casablanca. In a region inhabited since the Paleolithic Era over 300,000 years ago, the first Moroccan s ...
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Vertebra
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic irregular bone whose complex structure is composed primarily of bone, and secondarily of hyaline cartilage. They show variation in the proportion contributed by these two tissue types; such variations correlate on one hand with the cerebral/caudal rank (i.e., location within the backbone), and on the other with phylogenetic differences among the vertebrate taxa. The basic configuration of a vertebra varies, but the bone is its ''body'', with the central part of the body constituting the ''centrum''. The upper (closer to) and lower (further from), respectively, the cranium and its central nervous system surfaces of the vertebra body support attachment to the intervertebral discs. The posterior part of a vertebra forms a vertebral arch ...
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Appendicular Skeleton
The appendicular skeleton is the portion of the skeleton of vertebrates consisting of the bones that support the appendages. There are 126 bones. The appendicular skeleton includes the skeletal elements within the limbs, as well as supporting shoulder girdle and pelvic girdle.Skeleton
''Encyclopædia Britannica''. Updated 24 August 2014. The word appendicular is the adjective of the noun ''appendage'', which itself means a part that is joined to something larger.


The organization of the appendicular system

Of the 206 bones in the human skeleton, the appendicular skeleton comprises 126. Functionally it is involved in locomotion (lower limbs) of the

Foramina
In anatomy and osteology, a foramen (;Entry "foramen"
in
Merriam-Webster Online Dictionary
'. plural foramina, or foramens ) is an open hole that is present in extant or extinct s. Foramina inside the of typically allow ,



Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding age. Both age and stage bear the same name. As a unit of geologic time measure, the Cenomanian Age spans the time between 100.5 and 93.9 million years ago (Mya). In the geologic timescale, it is preceded by the Albian and is followed by the Turonian. The Upper Cenomanian starts around at 95 Mya. The Cenomanian is coeval with the Woodbinian of the regional timescale of the Gulf of Mexico and the early part of the Eaglefordian of the regional timescale of the East Coast of the United States. At the end of the Cenomanian, an anoxic event took place, called the Cenomanian-Turonian boundary event or the "Bonarelli event", that is associated with a minor extinction event for marine spec ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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