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Tryblidiida
Tryblidiida is a taxon of monoplacophoran molluscans containing the only extant representatives: 37 species are still alive today, inhabiting the ocean at depths of between . History of discoveries The first captured living monoplacophoran was ''Veleropilina zografi'' in 1896, but at that time it was described as if it were an archaeogastropod, a true limpet, mainly because of its patelliform (limpet-like) shell. This species was finally revealed to be monoplacophoran 87 years later, in 1983. In April 1952, a living specimen was collected from deep depths in the Middle America Trench off Costa Rica's Pacific coast. In 1957 that species was described and named ''Neopilina galatheae'' by its discoverer, Danish biologist Henning Mourier Lemche (1904–1977). An expert in the field has called this discovery "one of the greatest sensations in the [twentieth] century."Schwabe, above, at p. 205. As of 2008, there were 31 living species known, discovered in waters from 200 meters in de ...
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Monoplacophora
Monoplacophora , meaning "bearing one plate", is a polyphyletic superclass of molluscs with a cap-like shell inhabiting deep sea environments . Extant representatives were not recognized as such until 1952; previously they were known only from the fossil record, and were thought to have become extinct 375 million years ago. Although the shell of many monoplacophorans is limpet-like in shape, they are not gastropods, nor do they have any close relation to gastropods. Definition Discussion about monoplacophorans is made difficult by the slippery definition of the taxon; some authors take it to refer to all non-gastropod mollusks with a single shell, or all single-shelled mollusks with serially repeated units; whereas other workers restrict the definition to cap-shaped forms, excluding spiral and other shapes of shell. The inclusion of the gastropod-like Bellerophontoidea within the group is also contentious. One attempt to resolve this confusion was to separate out the predom ...
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Henning Mourier Lemche
Henning Mourier Lemche (11 August 1904, Copenhagen – 4 August 1977) was a Danish zoologist. Henning Mourier Lemche was the son of Søren Jacobsen Lemche and Inge Lemche née Mourier. He was educated at the University of Copenhagen, where he received his doctorate in 1937. He married Inger Sodemann on 17 April 1932. They had three children. From 1924 to 1948, he worked in of the Laboratory of Zoology at the Royal Veterinary and Agricultural University located in Frederiksberg, Denmark. Starting in 1949, he worked in the Department of Zoology at the University of Copenhagen. He became Curator of Molluscs at the Zoological Museum in 1955, and Master Assistant in 1962. After 1948, he was a member of the International Commission on Zoological Nomenclature. He wrote ''Fra Molekyle til Menneske'' (1945) and many scientific papers. He was interested in insects as well as nudibranchs. Henning Mourier Lemche discovered the mollusc ''Neopilina galatheae'', one of the three last living s ...
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Neopilina
''Neopilina'' is a highly derived genus of modern monoplacophoran.Organisms, Genes and Evolution: Evolutionary Theory at the Crossroads ; Proceedings of the 7th International Senckenberg Conference. By Dieter Stefan Peters, Michael Weingarten. Contributor: Dieter Stefan Peters. Published by Franz Steiner Verlag, 2000. , 9783515076593. 243 pages. Species * ''Neopilina bruuni'' Menzies, 1968 * ''Neopilina galatheae'' Lemche, 1957 * ''Neopilina rebainsi'' Moskalev, Starobogatov & Filatova, 1983 * ''Neopilina starobogatovi'' Ivanov & Moskalev, 2007 Phylogeny Some molecular results show that they fall within the polyplacophoran clade, although these results have been called into question. Fossil and morphological data show that they are rather derived and bear very little resemblance to an 'ancestral mollusc'. Anatomy Its anatomy led researchers to believe that the cephalopods evolved from the Monoplacophora. Its pair of preoral tentacles are considered homologous to those of gas ...
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Pleistocene
The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology. The name is a combination of Ancient Greek grc, label=none, πλεῖστος, pleīstos, most and grc, label=none, καινός, kainós (latinized as ), 'new'. At the end of the preceding Pliocene, the previously isolated North and South American continents were joined by the Isthmus of Panama, causing Great American Interchang ...
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Anus
The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, depending on the type of animal, includes: matter which the animal cannot digest, such as bones; Summary at food material after the nutrients have been extracted, for example cellulose or lignin; ingested matter which would be toxic if it remained in the digestive tract; and dead or excess gut bacteria and other endosymbionts. Amphibians, reptiles, and birds use the same orifice (known as the cloaca) for excreting liquid and solid wastes, for copulation and egg-laying. Monotreme mammals also have a cloaca, which is thought to be a feature inherited from the earliest amniotes via the therapsids. Marsupials have a single orifice for excreting both solids and liquids and, in females, a separate vagina for reproduction. Female placental mamm ...
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Chiton
Chitons () are marine molluscs of varying size in the class Polyplacophora (), formerly known as Amphineura. About 940 extant and 430 fossil species are recognized. They are also sometimes known as gumboots or sea cradles or coat-of-mail shells or suck-rocks, or more formally as loricates, polyplacophorans, and occasionally as polyplacophores. Chitons have a shell composed of eight separate shell plates or valves. These plates overlap slightly at the front and back edges, and yet articulate well with one another. Because of this, the shell provides protection at the same time as permitting the chiton to flex upward when needed for locomotion over uneven surfaces, and even allows the animal to curl up into a ball when dislodged from rocks. The shell plates are encircled by a skirt known as a girdle. Habitat Chitons live worldwide, from cold waters through to the tropics. They live on hard surfaces, such as on or under rocks, or in rock crevices. Some species live quite hi ...
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Annelids
The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecologies – some in marine environments as distinct as tidal zones and hydrothermal vents, others in fresh water, and yet others in moist terrestrial environments. The Annelids are bilaterally symmetrical, triploblastic, coelomate, invertebrate organisms. They also have parapodia for locomotion. Most textbooks still use the traditional division into polychaetes (almost all marine), oligochaetes (which include earthworms) and leech-like species. Cladistic research since 1997 has radically changed this scheme, viewing leeches as a sub-group of oligochaetes and oligochaetes as a sub-group of polychaetes. In addition, the Pogonophora, Echiura and Sipuncula, previously regarded as separate phyla, are now regarded as sub-groups of polychae ...
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Mollusks
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8 taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastropods ...
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Transitional Fossil
A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation. Because of the incompleteness of the fossil record, there is usually no way to know exactly how close a transitional fossil is to the point of divergence. Therefore, it cannot be assumed that transitional fossils are direct ancestors of more recent groups, though they are frequently used as models for such ancestors. In 1859, when Charles Darwin's ''On the Origin of Species'' was first published, the fossil record was poorly known. Darwin described the perceived lack of transitional fossils as "the most obvious and gravest objection which can be ...
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Segmentation (biology)
Segmentation in biology is the division of some animal and plant body plans into a series of repetitive segments. This article focuses on the segmentation of animal body plans, specifically using the examples of the taxa Arthropoda, Chordata, and Annelida. These three groups form segments by using a "growth zone" to direct and define the segments. While all three have a generally segmented body plan and use a growth zone, they use different mechanisms for generating this patterning. Even within these groups, different organisms have different mechanisms for segmenting the body. Segmentation of the body plan is important for allowing free movement and development of certain body parts. It also allows for regeneration in specific individuals. Definition Segmentation is a difficult process to satisfactorily define. Many taxa (for example the molluscs) have some form of serial repetition in their units but are not conventionally thought of as segmented. Segmented animals are tho ...
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Metamerism (biology)
In biology, metamerism is the phenomenon of having a linear series of body segments fundamentally similar in structure, though not all such structures are entirely alike in any single life form because some of them perform special functions. In animals, metameric segments are referred to as somites or metameres. In plants, they are referred to as metamers or, more concretely, phytomers. In animals In animals, zoologists define metamery as a mesodermal event resulting in serial repetition of unit subdivisions of ectoderm and mesoderm products. Endoderm is not involved in metamery. Segmentation is not the same concept as metamerism: segmentation can be confined only to ectodermally derived tissue, e.g., in the Cestoda tapeworms. Metamerism is far more important biologically since it results in metameres - also called somites - that play a critical role in advanced locomotion. One can divide metamerism into two main categories: * homonomous metamery is a strict serial succes ...
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Apex (mollusc)
In anatomy, an apex (adjectival form: apical) is part of the shell of a mollusk. The apex is the pointed tip (the oldest part) of the shell of a gastropod, scaphopod, or cephalopod. The apex is used in end-blown conches. Gastropods The word "apex" is most often used to mean the tip of the spire of the shell of a gastropod. The apex is the first-formed, and therefore the oldest, part of the shell. To be more precise, the apex would usually be where the tip of the embryonic shell or protoconch is situated, if that is still present in the adult shell (often it is lost or eroded away). Coiled gastropod shells The phrase apical whorls, or protoconch, means the whorls that constitute the embryonic shell at the apex of the shell, especially when this is clearly distinguishable from the later whorls of the shell, otherwise known as the teleoconch. Comparison of the apical part and the whole shell of ''Otukaia kiheiziebisu'': File:Calliostoma kiheiziebisu apex.png File:Calliostoma k ...
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