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Tropidostoma
''Tropidostoma'' is a medium-sized herbivorous oudenodontid dicynodont therapsid that lived during the Late Permian (Lopingian) period in South Africa. The first ''Tropidostoma'' fossil was described by Harry Govier Seeley in 1889. Later two subspecies were identified. ''Tropidostoma'' fossils are an index fossil in a biozone of the Karoo Basin known as the ''Tropidostoma'' Assemblage Zone. This biozone is characterized by the presence of this species in association with another dicynodont species, '' Endothiodon uniseries''. History of discovery The first ''Tropidostoma'' fossil material was found during a field expedition in the Upper Permian-aged Teekloof Formation of the Beaufort Group. This material was later described by Seeley (1889) in a study in which he described two fossils which had been named ''Dicynodon microtrema'' and ''Tropidostoma dunni''. In 1915, several years after Seeley's death, the paleontologist, Robert Broom, reexamined the same material and discover ...
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Bulbasaurus
''Bulbasaurus'' (meaning "bulbous reptile") is an extinct genus of dicynodont that is known from the Lopingian epoch of the Late Permian period of what is now South Africa, containing the type and only species ''B. phylloxyron''. It was formerly considered as belonging to '' Tropidostoma''; however, due to numerous differences from ''Tropidostoma'' in terms of skull morphology and size, it has been reclassified the earliest known member of the family Geikiidae, and the only member of the group known from the ''Tropidostoma'' Assemblage Zone. Within the Geikiidae, it has been placed close to '' Aulacocephalodon'', although a more basal position is not implausible. ''Bulbasaurus'' was ostensibly not directly named after the Pokémon Bulbasaur, but rather after its nasal bosses, which are unusually bulbous among geikiids; however, the describers noted that the similarity in name "may not be entirely coincidental." Additionally, the specific name of the type species means "leaf ra ...
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Tropidostoma Assemblage Zone
The ''Tropidostoma'' Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the lower Teekloof Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. The thickest outcrops, reaching approximately , occur from east of Sutherland through to Beaufort West and Victoria West, to areas south of Graaff-Reinet. Its northernmost exposures occur west/north-west of Colesberg. The '' Tropidostoma'' Assemblage Zone is the fourth biozone of the Beaufort Group. The name of the biozone refers to '' Tropidostoma microtrema'', a herbivorous dicynodont therapsid. This biozone is characterized by the presence of this species in association with another dicynodont species, '' Endothiodon uniseries''. History The first fossils to be found in the Beaufort Group rocks that encompass the current eight biozones were discovered by Andrew Geddes Bain in 1856. However, it was not until ...
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Beaufort Group
The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably underlies the Stormberg Group. Based on stratigraphic position, lithostratigraphic and biostratigraphic correlations, palynological analyses, and other means of geological dating, the Beaufort Group rocks are considered to range between Middle Permian ( Wordian) to Early Triassic (Anisian) in age. Background During the period when sedimentation of the Beaufort Group rocks took place, the Ecca sea had retreated to the northeastern Karoo Basin. All sediment deposition at this time took place in a terrestrial, although in a predominantly fluvial or alluvial environment that was seasonally arid. This environment covered a vast area and deposition was influenced by a retroarc foreland basin. This foreland system was caused by crustal upli ...
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Dicynodonts
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers. Characteristics The dicynodont skull is highly specialised, light but strong, with the synapsid temporal openings at the rear of the skull ...
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Dicynodont
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers. Characteristics The dicynodont skull is highly specialised, light but strong, with the synapsid temporal openings at the rear of the sk ...
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List Of Therapsids
This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (''nomina dubia''), or were not formally published ('' nomina nuda''), as well as junior synonyms of more established names, and genera that are no longer considered therapsids. The list currently contains 510 generic names. Naming conventions and terminology Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include: * Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior syno ...
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Oudenodon
''Oudenodon'' is an extinct genus of dicynodont. It was common throughout southern Africa during the Late Permian. Several species of ''Oudenodon'' are known. Both ''O. bainii'', the type species, and ''O. grandis'' are known from South Africa. Specimens of ''O. luangwensis'' have been found from Zambia. One species, ''O. sakamenensis'', is the only Permian therapsid yet known from Madagascar.Mazin, J. M. and King, G. M. (1991). The first dicynodont from the Late Permian of Malagasy. ''Palaeontology'' 34:837–842. It is the type genus of the family Oudenodontidae, which includes members such as '' Tropidostoma''. See also * List of therapsids References The main groups of non-mammalian synapsids at Mikko's Phylogeny Archive External links ''Oudenodon''in the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database (PBDB) or ...
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Endothiodon
''Endothiodon'' (/ɛndoʊθiːoʊdɔːn/ "inner tooth" from Greek endothi (ἔνδοθῐ), "within", and odon (ὀδών), "tooth", most likely named for the characteristic of the teeth being placed internally to the maxillaBoos A. S., Schultz C. L., Vega C. S., Aumond J. J. "On the presence of the Late Permian dicynodont ''Endothiodon'' in Brazil" Palaeontology 56:4, 837-848 (2013)) is an extinct genus of large dicynodont from the Late Permian. Like other dicynodonts, ''Endothiodon'' was an herbivore, but it lacked the two tusks that characterized most other dicynodonts. The anterior portion of the upper and lower jaw are curved upward, creating a distinct beak that is thought to have allowed them to be specialized grazers.Latimer E. M., Gow C. E., Rubidge B. S. "Dentition and feeding niche of ''Endothiodon'' (Synapsida;Anomodontia)" Palaeontologia Africana 32, 75-82 (1995) ''Endothiodon'' was widespread and is found across the southern region of what was then a single large co ...
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Sexual Dimorphism
Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most animals and some plants. Differences may include secondary sex characteristics, size, weight, colour, markings, or behavioural or cognitive traits. These differences may be subtle or exaggerated and may be subjected to sexual selection and natural selection. The opposite of dimorphism is ''monomorphism'', which is when both biological sexes are phenotypically indistinguishable from each other. Overview Ornamentation and coloration Common and easily identified types of dimorphism consist of ornamentation and coloration, though not always apparent. A difference in coloration of sexes within a given species is called sexual dichromatism, which is commonly seen in many species of birds and reptiles. Sexual selection leads to the exaggerated ...
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Tusk
Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share common features such as extra-oral position, growth pattern, composition and structure, and lack of contribution to ingestion. Tusks are thought to have adapted to the extra-oral environments, like dry or aquatic or arctic. In most tusked species both the males and the females have tusks although the males' are larger. Most mammals with tusks have a pair of them growing out from either side of the mouth. Tusks are generally curved and have a smooth, continuous surface. The male narwhal's straight single helical tusk, which usually grows out from the left of the mouth, is an exception to the typical features of tusks described above. Continuous growth of tusks is enabled by formative tissues in the apical openings of the roots of the teeth ...
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Maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxillary bones are fused at the intermaxillary suture, forming the anterior nasal spine. This is similar to the mandible (lower jaw), which is also a fusion of two mandibular bones at the mandibular symphysis. The mandible is the movable part of the jaw. Structure In humans, the maxilla consists of: * The body of the maxilla * Four processes ** the zygomatic process ** the frontal process of maxilla ** the alveolar process ** the palatine process * three surfaces – anterior, posterior, medial * the Infraorbital foramen * the maxillary sinus * the incisive foramen Articulations Each maxilla articulates with nine bones: * two of the cranium: the frontal and ethmoid * seven of the face: the nasal, zygomatic, lacrimal ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between org ...
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