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Tokosaurus
''Macroleter'' is an extinct genus of nycteroleterid parareptile which existed in Oklahoma and Russia during the upper Permian period. It was a quite generalized primitive reptile, in many ways resembling their amphibian ancestors. It was first named by paleontologists Tverdochlebova and Ivachnenko in 1984. According to classification by Michel Laurin and Robert R. Reisz, the genus is a parareptile, belonging to the same branch as Millerettidae, Procolophonidae and other generalized anapsid reptiles. The type species is ''Macroleter poezicus'' from Upper Permian of Russia. ''Macroleter'' had an 8 cm skull, and an overall length of 75 cm. It was generally lizard-like in build with a rather flat and broad skull. The teeth were small and pointy, indication it predominantly hunted insects and other small invertebrates. ''Seymouria agilis'' (Olson, 1980) that is known from only one specimen (holotype UCMP 143 277) was originally thought to be a reptile-like amphibian a ...
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Tokosaurus BW
''Macroleter'' is an extinct genus of nycteroleterid parareptile which existed in Oklahoma and Russia during the upper Permian period. It was a quite generalized primitive reptile, in many ways resembling their amphibian ancestors. It was first named by paleontologists Tverdochlebova and Ivachnenko in 1984 in paleontology, 1984. According to classification by Michel Laurin and Robert R. Reisz, the genus is a Parareptilia, parareptile, belonging to the same branch as Millerettidae, Procolophonidae and other generalized anapsid reptiles. The type species is ''Macroleter poezicus'' from Upper Permian of Russia. ''Macroleter'' had an 8 cm skull, and an overall length of 75 cm. It was generally lizard-like in build with a rather flat and broad skull. The teeth were small and pointy, indication it predominantly hunted insects and other small invertebrates. ''Seymouria agilis'' (Olson, 1980) that is known from only one specimen (holotype UCMP 143 277) was originally thought ...
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Nycteroleterid
Nycteroleteridae is a family of procolophonian parareptilians (extinct early reptiles) from the Middle to Late Permian of Russia and North America. They are sometimes classified as a sister group to pareiasaurids (but see ''Classification)''. The group includes the genera '' Macroleter'', '' Bashkyroleter'', ''"Bashkyroleter" mesensis'', '' Nycteroleter'', '' Emeroleter'', and probably '' Rhipaeosaurus''. They were carnivorous, and occasionally ate insects. The group was most common in European Russia, with only a few fossils in North America. One fossil has also been found in Africa, but this is the only one from Gondwana. Classification Nycteroleteridae is sometimes considered a sister group to the Pareiasauridae, but Bayesian inference suggests that it was in fact paraphyletic, with '' Rhipaeosaurus'' a basal member of the Pareiasauridae and other members of the Nycteroleteridae as outgroups. This is supported by the appearance of ''Rhipaeosaurus skull and teeth - ...
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Late Permian
Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, a concept in econometrics Music * Late (album), ''Late'' (album), a 2000 album by The 77s * Late!, a pseudonym used by Dave Grohl on his ''Pocketwatch (album), Pocketwatch'' album * Late (rapper), an underground rapper from Wolverhampton * Late (song), "Late" (song), a song by Blue Angel * "Late", a song by Kanye West from ''Late Registration'' Other * Late (Tonga), an uninhabited volcanic island southwest of Vavau in the kingdom of Tonga * Late (The Handmaid's Tale), "Late" (''The Handmaid's Tale''), a television episode * LaTe, Laivateollisuus, Oy Laivateollisuus Ab, a defunct shipbuilding company * Late may refer to a person who is Dead See also

* * * ''Lates'', a genus of fish in the lates perch family * Later (other) ...
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Millerettidae
Millerettidae is an extinct family of parareptiles from the Middle Permian to the Late Permian period (Capitanian - Changhsingian stages) of South Africa. The millerettids were small insectivores and probably resembled modern lizards in appearance and lifestyle. The following cladogram shows the phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... position of the Millerettidae, from Ruta ''et al.'', 2011. References External linksBerkeley University information. Permian reptiles Prehistoric reptiles of Africa Guadalupian first appearances Lopingian extinctions Prehistoric reptile families Parareptiles {{Permian-reptile-stub ...
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Permian Reptiles
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids (reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their amphibian ...
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Chickasha Formation
The Chickasha Formation is a geologic formation in Oklahoma. It preserves fossils dating back to the Permian period. See also * List of fossiliferous stratigraphic units in Oklahoma * Paleontology in Oklahoma Paleontology in Oklahoma refers to paleontological research occurring within or conducted by people from the U.S. state of Oklahoma. Oklahoma has a rich fossil record spanning all three eras of the Phanerozoic Eon. Oklahoma is the best source of ... References * Permian geology of Oklahoma {{Permian-stub ...
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Skeleton
A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside the body, and the hydroskeleton, a flexible internal skeleton supported by fluid pressure. Vertebrates are animals with a vertebral column, and their skeletons are typically composed of bone and cartilage. Invertebrates are animals that lack a vertebral column. The skeletons of invertebrates vary, including hard exoskeleton shells, plated endoskeletons, or Sponge spicule, spicules. Cartilage is a rigid connective tissue that is found in the skeletal systems of vertebrates and invertebrates. Etymology The term ''skeleton'' comes . ''Sceleton'' is an archaic form of the word. Classification Skeletons can be defined by several attributes. Solid skeletons consist of hard substances, such as bone, cartilage, or cuticle. These can be further ...
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Reptiliomorpha
Reptiliomorpha (meaning reptile-shaped; in PhyloCode known as ''Pan-Amniota'') is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians ( lissamphibians). It was defined by Michel Laurin (2001) and Vallin and Laurin (2004) as the largest clade that includes ''Homo sapiens'', but not ''Ascaphus truei'' (tailed frog). Laurin and Reisz (2020) defined Pan-Amniota as the largest total clade containing ''Homo sapiens'', but not ''Pipa pipa'', '' Caecilia tentaculata'', and '' Siren lacertina''. The informal variant of the name, "reptiliomorphs", is also occasionally used to refer to stem-amniotes, i.e. a grade of reptile-like tetrapods that are more closely related to amniotes than they are to lissamphibians, but are not amniotes themselves; the name is used in this meaning e.g. by Ruta, Coates and Quicke (2003). An alternative name, "Anthracosauria", is also commonly used for the group, but is confusing ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Upper Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids (reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their amp ...
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Anapsid
An anapsid is an amniote whose skull lacks one or more skull openings (fenestra, or fossae) near the temples. Traditionally, the Anapsida are the most primitive subclass of amniotes, the ancestral stock from which Synapsida and Diapsida evolved, making anapsids paraphyletic. It is however doubtful that all anapsids lack temporal fenestra as a primitive trait, and that all the groups traditionally seen as anapsids truly lacked fenestra. Anapsids and the turtles While "anapsid reptiles" or "anapsida" were traditionally spoken of as if they were a monophyletic group, it has been suggested that several groups of reptiles that had anapsid skulls might be only distantly related. Scientists still debate the exact relationship between the basal (original) reptiles that first appeared in the late Carboniferous, the various Permian reptiles that had anapsid skulls, and the Testudines (turtles, tortoises, and terrapins). However, it was later suggested that the anapsid-like turtle skull is ...
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