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Titanotaria
''Titanotaria'' is a genus of late, basal Odobenidae, walrus from the Miocene of Orange County, California. Unlike much later odobenids, it lacked tusks. ''Titanotaria'' is known from an almost complete specimen which serves as the holotype for the only recognized species, ''Titanotaria orangensis'', it is the best preserved fossil walrus currently known. History and naming Although the holotype specimen (OCPC 11141) of ''Titanotaria'' had been discovered in 1993 and represents one of the most complete fossil walrus known, little attention was given to the material for over 20 years. The first mention of the fossils in peer-reviewed literature came in 2017 with Barboza and colleagues publishing a faunal list of the Oso Member of the Capistrano Formation, where ''Titanotaria'' had been found. Specifically, the fossilized bones were collected from the town of Lake Forest, California, Lake Forest, Orange County, California, during the construction of the Saddleback Church. A full des ...
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Titanotaria Dorsal
''Titanotaria'' is a genus of late, basal walrus from the Miocene of Orange County, California. Unlike much later odobenids, it lacked tusks. ''Titanotaria'' is known from an almost complete specimen which serves as the holotype for the only recognized species, ''Titanotaria orangensis'', it is the best preserved fossil walrus currently known. History and naming Although the holotype specimen (OCPC 11141) of ''Titanotaria'' had been discovered in 1993 and represents one of the most complete fossil walrus known, little attention was given to the material for over 20 years. The first mention of the fossils in peer-reviewed literature came in 2017 with Barboza and colleagues publishing a faunal list of the Oso Member of the Capistrano Formation, where ''Titanotaria'' had been found. Specifically, the fossilized bones were collected from the town of Lake Forest, Orange County, California, during the construction of the Saddleback Church. A full description followed a year after its ...
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Capistrano Formation
The Capistrano Formation is a geologic formation in coastal southern Orange County, California. It preserves fossils dating back to the late Miocene to early Pliocene, with the Oso Member representing a near-shore environment. Fifty-nine species and varieties of foraminifera are recognized from the Capistrano Formation alongside a diverse array of marine mammals including up to five species of walrus. Geography The Capistrano Formation, named for the town of San Juan Capistrano, is located in southern California, specifically the northern extent of the Peninsular Ranges, which stretch from the Los Angeles Basin to Baja California. It crops out along the coast from Dana Point to San Clemente, and inland for seven miles. Geology and Stratigraphy The Capistrano Formation is a heterogenous marine formation that can be differentiated into two distinct but adjacent units. One of these units is the Oso Member, which is composed of arkosic sandstone and preserves a nearshore environm ...
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Osodobenus
''Osodobenus'' is an extinct genus of walrus from the Miocene to Pliocene of California. ''Osodobenus'' may have been the first tusked walrus and shows several adaptations that suggest it was a suction feeder, possibly even a benthic feeder like modern species. Three skulls are known showing pronounced sexual dimorphism, with the female lacking the same tusks as the male. Only a single species, ''Osodobenus eodon'', is currently recognized. Discovery and naming ''Osodobenus'' is known from three specimens including an adult male, adult female and a juvenile specimen preserving skulls and some postcranial material. All the material was collected from the Late Miocene to Early Pliocene (Messinian to Zanclean) Capistrano Formation, Orange County, California alongside the remains of several other early odobenids. In 2020, Biewer and colleagues published a detailed description of the material, establishing ''Osodobenus'' as a new genus while also erecting two new species of ''Pontolis ...
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Odobenidae
Odobenidae is a family of pinnipeds. The only living species is the walrus (''Odobenus rosmarus''). In the past, however, the group was much more diverse, and includes more than a dozen fossil genera. Taxonomy All genera, except ''Walrus, Odobenus'', are extinct. *†''Archaeodobenus'' *†''Prototaria'' *†''Proneotherium'' *†''Nanodobenus'' *†''Neotherium'' *†''Imagotaria'' *†''Kamtschatarctos'' *†''Pelagiarctos'' *†''Pontolis'' *†''Pseudotaria'' *†''Titanotaria'' *Clade Neodobenia **†''Gomphotaria'' **Subfamily Dusignathinae ***†''Dusignathus'' **Subfamily Odobeninae ***†''Aivukus'' ***†''Ontocetus'' ***†''Pliopedia'' ***†''Protodobenus'' ***†''Valenictus'' ***''Walrus, Odobenus'' In re-analyzing ''Pelagiarctos'', Boessenecker et al. (2013) proposed the phylogenetic relationships of Odobenidae as follows (this analysis excluded ''Archaeodobenus, Titanotaria, Nanodobenus,'' and ''Pliopedia;'' and included ''Enaliarctos, Pteronarctos, Allodesmus, ...
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Neodobenia
Odobenidae is a family of pinnipeds. The only living species is the walrus (''Odobenus rosmarus''). In the past, however, the group was much more diverse, and includes more than a dozen fossil genera. Taxonomy All genera, except ''Odobenus'', are extinct. *†''Archaeodobenus'' *†''Prototaria'' *†''Proneotherium'' *†''Nanodobenus'' *†''Neotherium'' *†''Imagotaria'' *†''Kamtschatarctos'' *†''Pelagiarctos'' *†''Pontolis'' *†'' Pseudotaria'' *†''Titanotaria'' *Clade Neodobenia **†''Gomphotaria'' **Subfamily Dusignathinae ***†'' Dusignathus'' **Subfamily Odobeninae ***†''Aivukus'' ***†''Ontocetus'' ***†''Pliopedia'' ***†'' Protodobenus'' ***†'' Valenictus'' ***''Odobenus The walrus (''Odobenus rosmarus'') is a large flippered marine mammal with a discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. The walrus is the only living species in the fa ...'' In re-analyzing ''Pelagiar ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Archaeodobenus
''Archaeodobenus'' is an extinct genus of pinniped that lived during the Late Miocene of what is now Japan. It belonged to the Odobenidae family, which is today only represented by the walrus, but was much more diverse in the past, containing at least 16 genera. Unlike the modern walrus, ''Archaeodobenus'' did not have tusks but instead had canines of moderate size, and looked more like a sea lion. Discovery The first known specimen was collected in 1977 from the Ichibangawa Formation in Tobetsu Town on the island of Hokkaido. The specimen consists of a partial skull, vertebrae, and limb bones, and was made the holotype specimen of the new genus and species ''A. akamatsui'' by the Japanese palaeontologists Yoshihiro Tanaka and Naoki Kohno in 2015. The generic name consists of ''archaio-'', the Greek word for ancient, and the generic name of the walrus, ''Odobenus''; in full, "ancient walrus." The specific name honors Morio Akamatsu, a curator of the Hokkaido Museum. The ho ...
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Kamtschatarctos
''Kamtschatarctos'' is an extinct genus of pinniped that lived approximately 15.97 to 11.608 mya during the Early Miocene in the Kavran-Ukhtolok Bay of Russia's Kamchatka Peninsula. It belonged to the family Odobenidae, the only extant species of which is the walrus The walrus (''Odobenus rosmarus'') is a large pinniped, flippered marine mammal with a discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. The walrus is the only living species in .... Discovery ''Kamtschatarctos sinelnikovae'' is known from a partially complete skeleton, discovered in the Etolon Formation in Russia. Taxonomy ''Kamtschatarctos'' is a relatively basal species of odobenid. References {{Taxonbar, from=Q48465840 Pinnipeds of Europe Odobenids Prehistoric pinnipeds Prehistoric carnivoran genera ...
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Proneotherium
''Proneotherium'' is an extinct genus of pinniped that lived approximately 20.43 to 15.97 mya during the Early Miocene in what is now Oregon, U.S. It belonged to the family Odobenidae, the only extant species of which is the walrus. Discovery Four specimens of ''Proneotherium'' were discovered in the Astoria Formation of Lincoln County, Oregon, U.S. These specimens included cranial and postcranial remains and are all believed to be adult male animals. Description ''Proneotherium'' were more similar in appearance to modern fur seals and sea lions than walruses. They lacked the long tusks of walruses, and were more slender and smaller than their modern relatives. Autapomorphies of ''Proneotherium'' fossils include a continuous, horizontal crest connecting the mastoid and paroccipital processes of the skull. The teeth are also less secodont in function and appearance than more basal species. This likely represents the slow shift from a piscivorous diet to the mollusk-based d ...
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Prototaria
''Prototaria'' is an extinct genus of pinniped that lived approximately 15.97 to 13.65 mya during the Middle Miocene in what is now Japan. It belonged to the family Odobenidae, the only extant species of which is the walrus. Members of the genus ''Prototaria'' are believed to be the most basal imagotariine pinnipeds. Unlike their closest living relative, the walrus, members of ''Prototaria'' were primarily piscivorous carnivores. Description ''Prototaria'' were more similar in appearance to modern fur seals and sea lions than walruses. They did not have long tusks as walruses do, and were more slender. Fossils ascribed to ''Prototaria'' exhibit some autapomorphies, including: a large antorbital process and orbit, and a relatively narrow intertemporal region of the skull. ''P. planicephala'' has a wider rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mout ...
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Pelvis
The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The pelvic region of the trunk includes the bony pelvis, the pelvic cavity (the space enclosed by the bony pelvis), the pelvic floor, below the pelvic cavity, and the perineum, below the pelvic floor. The pelvic skeleton is formed in the area of the back, by the sacrum and the coccyx and anteriorly and to the left and right sides, by a pair of hip bones. The two hip bones connect the spine with the lower limbs. They are attached to the sacrum posteriorly, connected to each other anteriorly, and joined with the two femurs at the hip joints. The gap enclosed by the bony pelvis, called the pelvic cavity, is the section of the body underneath the abdomen and mainly consists of the reproductive organs (sex organs) and the rectum, while the pelvic f ...
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