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Thyreophora
Thyreophora ("shield bearers", often known simply as "armored dinosaurs") is a group of armored ornithischian dinosaurs that lived from the Early Jurassic until the end of the Cretaceous. Thyreophorans are characterized by the presence of body armor lined up in longitudinal rows along the body. Primitive forms had simple, low, keeled scutes or osteoderms, whereas more derived forms developed more elaborate structures including spikes and plates. Most thyreophorans were herbivorous and had relatively small brains for their body size. Thyreophora includes various subgroups, including the suborders Ankylosauria and Stegosauria. In both the suborders, the forelimbs were much shorter than the hindlimbs, particularly in stegosaurs. The clade has been defined as the group consisting of all species more closely related to ''Ankylosaurus'' than to ''Triceratops''. Thyreophora is the sister group of Cerapoda within Genasauria. Groups of thyreophorans Basal thyreophorans Basal thyreop ...
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Genasauria
Genasauria is a clade of extinct beaked, primarily herbivorous dinosaurs. Paleontologist Paul Sereno first named Genasauria in 1986. The name Genasauria is derived from the Latin word ''gena'' meaning ‘cheek’ and the Greek word ''saúra'' (σαύρα) meaning ‘lizard.’ It is hypothesized that Genasauria had diverged from ''Lesothosaurus'' by the Early Jurassic. Cranial features that characterize Genasauria include a medial offset of the maxillary dentition, a sprout-shaped mandibular symphysis, moderately sized coronoid process, and an edentulous (without teeth) anterior portion of the premaxilla. A distinguishing postcranial feature of Genasauria is a pubic peduncle of the ilium that is less robust than the ischial peduncle. Genasauria is commonly divided into Neornithischia and Thyreophora. Neornithischia is characterized by asymmetrical distributions of enamel covering the crowns of the cheek teeth, an open acetabulum, and a laterally protruding ischial peduncle of the ...
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Emausaurus
''Emausaurus'' is a genus of thyreophoran or armored dinosaur from the Early Jurassic (Early Toarcian). Its fossils have been found in Mecklenburg-Vorpommern, northern Germany. ''Emausaurus'' is the only known Toarcian thyreophoran, as well as the only dinosaur from the zone of the same age with a formal name. Discovery and naming The type and only species, ''E. ernsti'', was named and described by Harmut Haubold in 1990. The generic name is composed of an acronym of Ernst Moritz Arndt University of Greifswald and the Greek ''/'' (lizard). The specific name is derived from the name of geologist Werner Ernst, who acquired the holotype specimen, SGWG 85, in the summer of 1963 from foreman Werner Wollin at a loampit near Grimmen, in strata dating from the Toarcian. It is known by the right side of the skull, the right lower jaw, caudal vertebrae, neural arches, a radius, a metatarsal, a claw, fragments of ribs, scutes, and plates, known as EMAU SGWG 85. Description ''Emausaurus ...
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Stegosaurus Stenops
''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been found in the western United States and in Portugal, where they are found in Kimmeridgian- to early Tithonian-aged strata, dating to between 155 and 145 million years ago. Of the species that have been classified in the upper Morrison Formation of the western US, only three are universally recognized: ''S. stenops'', ''S. ungulatus'' and ''S. sulcatus''. The remains of over 80 individual animals of this genus have been found. ''Stegosaurus'' would have lived alongside dinosaurs such as ''Apatosaurus'', ''Diplodocus'', ''Brachiosaurus'', ''Ceratosaurus'', and ''Allosaurus''; the latter two may have preyed on it. They were large, heavily built, herbivorous quadrupeds with rounded backs, short fore limbs, long hind limbs, and tails held high ...
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Scelidosaurus
''Scelidosaurus'' (; with the intended meaning of "limb lizard", from Greek / meaning 'rib of beef' and ''sauros''/ meaning 'lizard')Liddell & Scott (1980). Greek-English Lexicon, Abridged Edition. Oxford University Press, Oxford, UK. is a genus of herbivorous armoured ornithischian dinosaur from the Jurassic of the British Isles. ''Scelidosaurus'' lived during the Early Jurassic Period, during the Sinemurian to Pliensbachian stages around 191 million years ago. This genus and related genera at the time lived on the supercontinent Laurasia. Its fossils have been found in the Charmouth Mudstone Formation near Charmouth in Dorset, England, and these fossils are known for their excellent preservation. ''Scelidosaurus'' has been called the earliest complete dinosaur.Norman, David (2001). "''Scelidosaurus'', the earliest complete dinosaur" in ''The Armored Dinosaurs'', pp 3-24. Bloomington: Indiana University Press. . It is the most completely known dinosaur of the British Isle ...
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Laquintasaura
''Laquintasaura'' is a genus of Venezuelan Ornithischia, ornithischian dinosaur containing only the type species ''Laquintasaura venezuelae''. The species was the first dinosaur to have been identified from Venezuela. It is known from extensive remains, all from a singular bonebed locality which has been sampled for specimen blocks over the course of several expeditions, largely led by Marcelo R Sánchez-Villagra. A small, very primitive animal, it is known for its distinct dental anatomy and for being one of the earliest and most primitive ornithischians in the fossil record. Taxonomic uncertainty has led to conflicting theories that it is either as the base of Ornithischia or at the base of the subgroup Thyreophora. In either model, its dating to around 200 million years ago, at the start of the Jurassic, existence in equatorial latitudes, and primitive nature make it a key view into early ornithischian evolution. It's thought that ''Laquintasaura'' would've lived in groups and ...
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Ankylosauria
Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are known to have first appeared in the Middle Jurassic, and persisted until the end of the Cretaceous Period. The two main families of Ankylosaurs, Nodosauridae and Ankylosauridae are primarily known from the Northern Hemisphere, but the more basal Parankylosauria are known from southern Gondwana during the Cretaceous. Ankylosauria was first named by Henry Fairfield Osborn in 1923.Osborn, H. F. (1923). "Two Lower Cretaceous dinosaurs of Mongolia." ''American Museum Novitates'', 95: 1–1/ref> In the Linnaean classification system, the group is usually considered either a suborder or an infraorder. It is contained within the group Thyreophora, which also includes the stegosaurs, armored dinosaurs known for their combination of plate ...
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Yuxisaurus
''Yuxisaurus'' (meaning "Yuxi lizard") is an extinct genus of basal thyreophoran dinosaur from the Early Jurassic (Sinemurian-Toarcian) Fengjiahe Formation of southwestern China. The type species is ''Yuxisaurus kopchicki.'' Discovery and naming ''Yuxisaurus'' was first named in a ''bioRxiv'' preprint released in late 2021, but was not considered valid, due to having not been validly published. The taxon was formally published shortly thereafter, in March 2022. The holotype of the taxon, CVEB 21701, represents a partial skeleton including parts of the skull, several vertebrae, elements of the limbs, and at least 120 osteoderms. The generic name, "''Yuxisaurus''," combines a reference to the type locality in Yuxi, Yunnan Province, China, with the Greek "sauros," meaning "lizard." The specific name, "''kopchicki''," honors the biologist John J. Kopchick. Classification ''Yuxisaurus'' was a basal thyreophoran. Depending on the dataset, it may be the sister taxon to the Ger ...
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Ankylosaurus
''Ankylosaurus'' is a genus of armored dinosaur. Its fossils have been found in geological formations dating to the very end of the Cretaceous Period, about 68–66 million years ago, in western North America, making it among the last of the non-avian dinosaurs. It was named by Barnum Brown in 1908; it is monotypic, containing only ''A. magniventris''. The generic name means "fused" or "bent lizard", and the specific name means "great belly". A handful of specimens have been excavated to date, but a complete skeleton has not been discovered. Though other members of Ankylosauria are represented by more extensive fossil material, ''Ankylosaurus'' is often considered the archetypal member of its group, despite having some unusual features. Possibly the largest-known ankylosaurid, ''Ankylosaurus'' is estimated to have been between long and to have weighed between . It was quadrupedal, with a broad, robust body. It had a wide, low skull, with two horns pointing backward from the b ...
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Jakapil
''Jakapil'' (meaning "shield bearer" in Puelchean) is a genus of basal thyreophoran dinosaur from the Candeleros Formation of Argentina. The type species is ''Jakapil kaniukura''. Discovery and naming The holotype, MPCA-PV-630, is a partial skeleton including several osteoderms and a complete lower jaw, which were found on land owned by the Mariluan family in 2012 and excavated between 2014 and 2019/2020. The finds were prepared by L. Pazo and J. Kaluza. The generic name, "''Jakapil''", is derived from "Ja-Kapïl", a Puelchean word meaning "shield bearer". This is also the literal meaning of the clade name Thyreophora. The specific name, "''kaniukura''", means "crest stone" in Mapudungun, in reference to its uniquely deep jaw. Description ''Jakapil'' represents a novel morphotype among thyreophorans, including, among other things, the presence of a predentary bone (absent or cartilaginous in other basal thyreophorans), large, low osteoderms, and a bipedal stance, similar ...
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Stegosauria
Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Europe, Africa, South America and Asia. Their geographical origins are unclear; the earliest unequivocal stegosaurian, ''Huayangosaurus taibaii'', lived in China. Stegosaurians were armored dinosaurs (thyreophorans). Originally, they did not differ much from more primitive members of that group, being small, low-slung, running animals protected by armored scutes. An early evolutionary innovation was the development of spikes as defensive weapons. Later species, belonging to a subgroup called the Stegosauridae, became larger, and developed long hindlimbs that no longer allowed them to run. This increased the importance of active defence by the thagomizer, which could ward off even large predators because the tail was in a higher position, pointing ...
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Bienosaurus
''Bienosaurus'' (meaning "Bien's lizard") is a genus of thyreophoran dinosaur from the Lower Jurassic (probably Sinemurian) Lower Lufeng Formation in Yunnan Province in China. Discovery and species In 1938, Bian Meinian, known in the West as Mei Nien Bien, discovered a fragmentary dinosaur skull. In 2001, Dong Zhiming named and described the specimen as the type species ''Bienosaurus lufengensis''. The generic name honours Bian. The specific name refers to the Lufeng formation. ''Bienosaurus'' is based on holotype IVPP V15311 (in 2001 incorrectly given the inventory number IVPP V 9612). It consists of a partial right lower jaw with teeth and several cranial fragments. Phylogeny Dong in 2001 placed ''Bienosaurus'' in the Scelidosauridae, considering these to be part of the Ankylosauria. Later publications suggested a general position basal in the Thyreophora. In 2019 a study confirmed this, concluding ''Bienosaurus'' was a ''nomen dubium'', possibly identical to ''Tatisaur ...
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Scutellosaurus
''Scutellosaurus'' ( ) is a genus of thyreophoran ornithischian dinosaur that lived approximately 196 million years ago during the early part of the Jurassic Period in what is now Arizona, USA. It is classified in Thyreophora, the armoured dinosaurs; its closest relatives may have been ''Emausaurus'' and ''Scelidosaurus'', another armored dinosaur which was mainly a quadrupedal dinosaur, unlike bipedal ''Scutellosaurus''. It is one of the earliest representatives of the armored dinosaurs and the basalmost form discovered to date. ''Scutellosaurus'' was a small, lightly-built, ground-dwelling, herbivore, that could grow up to an estimated long. Etymology The genus name ''Scutellosaurus'' means "little-shielded lizard", and is derived from the Latin word "scutellum" meaning "little shield", and the Greek word "sauros" (σαύρα) meaning "lizard". The type and only valid species known today is ''Scutellosaurus lawleri''. The specific name honors David Lawler who collected the ...
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