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Thanatotheristes
''Thanatotheristes'' (meaning "reaper of death" according to the original authors; literally, "death harvester" in Greek language, Greek: , , "Death" and , , "harvester") is a genus of tyrannosaurid dinosaur from the Late Cretaceous of Laramidia, approximately 80.1-79.5 Ma. ''Thanatotheristes'' contains only one species, ''T. degrootorum''. Fossils of this taxon are found in the Foremost Formation of Alberta, Canada, coexisting with medium-sized Ceratopsidae, ceratopsids like ''Xenoceratops foremostensis'' and small Pachycephalosauridae, pachycephalosaurids like ''Colepiocephale lambei''. Discovery and naming The holotype specimen of ''Thanatotheristes degrootorum'' (TMP 2010.5.7) is based on a right maxilla, right jugal, right postorbital, right surangular, right quadrate, right laterosphenoid, left frontal, and both dentaries. The length of the skull has been approximated to be . It was smaller than the closely related ''Daspletosaurus'', but the holotype individual was not ...
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Thanatotheristes Size Comparison
''Thanatotheristes'' (meaning "reaper of death" according to the original authors; literally, "death harvester" in Greek: , , "Death" and , , "harvester") is a genus of tyrannosaurid dinosaur from the Late Cretaceous of Laramidia, approximately 80.1-79.5 Ma. ''Thanatotheristes'' contains only one species, ''T. degrootorum''. Fossils of this taxon are found in the Foremost Formation of Alberta, Canada, coexisting with medium-sized ceratopsids like ''Xenoceratops foremostensis'' and small pachycephalosaurids like ''Colepiocephale lambei''. Discovery and naming The holotype specimen of ''Thanatotheristes degrootorum'' (TMP 2010.5.7) is based on a right maxilla, right jugal, right postorbital, right surangular, right quadrate, right laterosphenoid, left frontal, and both dentaries. The length of the skull has been approximated to be . It was smaller than the closely related ''Daspletosaurus'', but the holotype individual was not osteologically mature at the time of death. A refer ...
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Daspletosaurus
''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 79.5 and 74 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three species. Fossils of the earlier type species, ''D. torosus'', have been found in Alberta, and fossils of a later second species, ''D. wilsoni,'' and third species, ''D. horneri'', have been found only in Montana. A possible fourth species, also from Alberta, awaits formal identification. The taxon ''Thanatotheristes'' has been suggested to represent a species of ''Daspletosaurus'', ''D. degrootorum'', but this has not been widely supported. ''Daspletosaurus'' is closely related to the much larger and more recent tyrannosaurid ''Tyrannosaurus rex''. Like most tyrannosaurids, ''Daspletosaurus'' was a multi-tonne bipedal predator equipped with dozens of large, sharp teeth. ''Daspletosaurus'' had the small forelimbs typical of tyrannosaurids, although ...
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Teratophoneus
''Teratophoneus'' ("monstrous murderer"; Greek: ''teras'', "monster" and ''phoneus'', "murderer") is a genus of tyrannosaurid dinosaur which lived during the late Cretaceous period (late Campanian age, about 77 to 76 million years ago) in what is now Utah, United States, containing a single known species, ''T. curriei''. It is known from an incomplete skull and postcranial skeleton recovered from the Kaiparowits Formation. It was specifically named ''T. curriei'' in honor of Philip J. Currie. Discovery and naming Fossils of ''Teratophoneus'' were first found in the Kaiparowits Formation of southern Utah. Later, fossils from the same formation were discovered and identified as the genus. Argon-argon radiometric dating indicates that the Kaiparowits Formation was deposited between 76.1 and 74.0 million years ago, during the Campanian stage of the Late Cretaceous period. This date means that ''Teratophoneus'' lived in the middle of the Campanian stage of the Late Cretaceous. ...
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Tarbosaurus
''Tarbosaurus'' ( ; meaning "alarming lizard") is a genus of tyrannosaurid dinosaur that flourished in Asia about 70 million years ago, at the end of the Late Cretaceous Period, considered to contain a single known species, ''Tarbosaurus bataar''. Fossils have been recovered from the Nemegt Formation of Mongolia, with more fragmentary remains found further afield in the Subashi Formation of China. Although many species have been named, modern paleontologists recognize only one, ''T. bataar'', as valid. Some experts see this species as an Asian representative of the North American genus ''Tyrannosaurus''; this would make the genus ''Tarbosaurus'' redundant. ''Tarbosaurus'' and ''Tyrannosaurus'', if not synonymous, are considered to be at least closely related genera. ''Alioramus'', also from Mongolia, has previously been thought by some authorities to be the closest relative of ''Tarbosaurus'', though this has since been disproven with the discovery of '' Qianzhousaurus'' and ...
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Lythronax
''Lythronax'' () is a genus of tyrannosaurid dinosaur that lived in North America around 81.9-81.5 million years ago during the Late Cretaceous period. The only known specimen was discovered in Utah in the Wahweap Formation of the Grand Staircase–Escalante National Monument in 2009, and it consists of a partial skull and skeleton. In 2013, it became the basis of the new genus and species ''Lythronax argestes''; the generic name ''Lythronax'' means "gore king", and the specific name ''argestes'' originates from the Greek poet Homer's name for the wind from the southwest, in reference to the specimen's geographic provenance in North America. Size estimates for ''Lythronax'' have ranged between in length, and between in weight. It was a heavily built tyrannosaurid, and as a member of that group, it would have had small, two-fingered forelimbs, strong hindlimbs, and a very robust skull. The rear part of the skull of ''Lythronax'' appears to have been very broad, with eye socket ...
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Bistahieversor
''Bistahieversor'' (meaning "Bistahi destroyer"), also known as the "Bisti Beast", is a genus of eutyrannosaurian tyrannosauroid dinosaur; the genus contains only a single known species, ''B. sealeyi'', described in 2010, from the Late Cretaceous of New Mexico. The holotype and a juvenile were found in the Hunter Wash Member of the Kirtland Formation, while other specimens came from the underlying Fossil Forest member of the Fruitland Formation. This dates ''Bistahieversor'' approximately 75.5 to 74.5 million years ago, found in sediments spanning a million years. Discovery and naming The first remains now attributed to ''Bistahieversor'', a partial skull and skeleton, were described in 1990 as a specimen of ''Aublysodon''. Additional remains, consisting of the incomplete skull and skeleton of a juvenile, were described in 1992. Another, complete, skull and partial skeleton were found in the Bisti/De-Na-Zin Wilderness of New Mexico in 1998, known colloquially as the "Bisti Beas ...
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Foremost Formation
The Foremost Formation is a formation (stratigraphy), stratigraphic unit of Late Cretaceous (Campanian) age that underlies much of southern Alberta, Canada. It was named for outcrops in Chin Coulee near the town of Foremost, Alberta, ForemostGlass, D.J. (editor) 1997. Lexicon of Canadian Stratigraphy, vol. 4, Western Canada including eastern British Columbia, Alberta, Saskatchewan and southern Manitoba. Canadian Society of Petroleum Geologists, Calgary; 1423 p. on CD. . and is known primarily for its dinosaur remains and other fossils.Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Cretaceous, North America)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press, pp. 574-588. . Geology The Foremost Formation is the basal unit of the Belly River Group (called the Judith River Group in the United States). It gradationally overlies the marine shales of the Pakowki Formation. It consists ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and Ant ...
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Colepiocephale Lambei
''Colepiocephale'' (meaning "knucklehead") is a genus of pachycephalosaurid dinosaur from Late Cretaceous (middle Campanian stage) deposits of Alberta, Canada. It was collected from the Foremost Formation (middle Campanian, 80–77.5 ma). The type species, ''C. lambei'', was originally described by Sternberg (in 1945 as ''Stegoceras lambei''C. M. Sternberg. 1945. Pachycephalosauridae proposed for dome-headed dinosaurs, Stegoceras lambei, n. sp., described. Journal of Paleontology 19(5):534-538), and later renamed by Sullivan in 2003. ''C. lambei'' is a domed pachycephalosaur characterized principally by the lack of a lateral and posteriosquamosal shelf, a steeply down-turned parietal, and the presence of two incipient nodes tucked under the posterior margin of the parietosquamosal border. See also * Timeline of pachycephalosaur research This timeline of pachycephalosaur research is a chronological listing of events in the history of paleontology focused on the pachycepha ...
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Albertosaurinae
Albertosaurines, or dinosaurs of the subfamily Albertosaurinae, lived in the Late Cretaceous of United States and Canada. The subfamily was first used by Philip J. Currie, Jørn H. Hurum and Karol Sabath as a group of tyrannosaurid dinosaurs. It was originally defined as "(''Albertosaurus'' + ''Gorgosaurus'')", including only the two genera. The group is sister clade to Tyrannosaurinae. In 2007, it was found that the group also contained '' Maleevosaurus'', often synonymized with ''Tarbosaurus''. However, this classification has not been accepted, and '' Maleevosaurus'' is still considered a juvenile ''Tarbosaurus'' or ''Tyrannosaurus''. Description Albertosaurines are large, lightly built tyrannosaurids. Compared to tyrannosaurines, they are lightly built, have shorter, flatter skulls, had shorter ilia, and had proportionally longer tibiae. Albertosaurines and tyrannosaurines share arms or about equal length, with the exception of ''Tarbosaurus'', which had sh ...
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Dinosaur Park Formation
The Dinosaur Park Formation is the uppermost member of the Belly River Group (also known as the Judith River Group), a major geologic unit in southern Alberta. It was deposited during the Campanian stage of the Late Cretaceous, between about 76.5 and 74.4 million years ago. It was deposited in alluvial and coastal plain environments, and it is bounded by the nonmarine Oldman Formation below it and the marine Bearpaw Formation above it.Eberth, D.A. 2005. The geology. In: Currie, P.J., and Koppelhus, E.B. (eds), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press: Bloomington and Indianapolis, p.54-82. . The Dinosaur Park Formation contains dense concentrations of dinosaur skeletons, both articulated and disarticulated, which are often found with preserved remains of soft tissues. Remains of other animals such as fish, turtles, and crocodilians, as well as plant remains, are also abundant. The formation has been named after Dinosaur Pr ...
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Tyrannosaurus
''Tyrannosaurus'' is a genus of large theropoda, theropod dinosaur. The species ''Tyrannosaurus rex'' (''rex'' meaning "king" in Latin), often called ''T. rex'' or colloquially ''T-Rex'', is one of the best represented theropods. ''Tyrannosaurus'' lived throughout what is now western North America, on what was then an island continent known as Laramidia. ''Tyrannosaurus'' had a much wider range than other Tyrannosauridae, tyrannosaurids. Fossils are found in a variety of geologic formation, rock formations dating to the Maastrichtian Age (geology), age of the Upper Cretaceous Period (geology), period, 68 to 66 mya (unit), million years ago. It was the last known member of the tyrannosaurids and among the last non-aves, avian dinosaurs to exist before the Cretaceous–Paleogene extinction event. Like other tyrannosaurids, ''Tyrannosaurus'' was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to its large and powerful hind limbs, the foreli ...
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