Temperate Herbaceous Clade
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Temperate Herbaceous Clade
The inverted repeat-lacking clade (IRLC) is a monophyletic clade of the flowering plant subfamily Faboideae (or Papilionaceae). Faboideae includes the majority of agriculturally-cultivated legumes. The name of this clade is informal and is not assumed to have any particular taxonomic rank like the names authorized by the ICBN or the ICPN. The clade is characterized by the loss of one of the two 25-kb inverted repeats in the plastid genome that are found in most land plants. It is consistently resolved in molecular phylogenies. The clade is predicted to have diverged from the other legume lineages 39.0±2.4 million years ago (in the Eocene). It includes several large, temperate genera such as '' Astragalus'' L., '' Hedysarum'' L., ''Medicago'' L., '' Oxytropis'' DC., ''Swainsona'' Salisb., and ''Trifolium'' L.. Description This clade is composed of five traditional tribes (Cicereae Alef., Fabeae Rchb., Galegeae Dumort., Hedysareae DC., Trifolieae Endl.) and several genera ...
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Galega Officinalis
''Galega officinalis'', commonly known as galega or goat's-rue, is an herbaceous plant in the subfamily Faboideae of the legume family Fabaceae. It is native to parts of northern Africa, western Asia and Europe, but is widely cultivated and naturalised elsewhere. The plant has been extensively cultivated as a forage crop, an ornamental, a bee plant, and as green manure. ''G. officinalis'' is rich in galegine, a substance with blood glucose-lowering activity and the foundation for the discovery of metformin, Italian fitch, a treatment for managing symptoms of diabetes mellitus. In ancient herbalism, goat's-rue was used as a diuretic. It can be poisonous to mammals, but is a food for various insects. Etymology The English name "goat's-rue" is a translation of the Latin ''Ruta capraria'', used for the plant in 1554 when it was considered to be related to ''Ruta graveolens'', or common rue. The Latin specific epithet ''officinalis'' refers to plants with some medicinal, culinary or ...
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ICBN
The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all those "traditionally treated as algae, fungi, or plants".. It was formerly called the ''International Code of Botanical Nomenclature'' (ICBN); the name was changed at the International Botanical Congress in Melbourne in July 2011 as part of the ''Melbourne Code''. which replaced the ''Vienna Code'' of 2005. The current version of the code is the ''Shenzhen Code'' adopted by the International Botanical Congress held in Shenzhen, China, in July 2017. As with previous codes, it took effect as soon as it was ratified by the congress (on 29 July 2017), but the documentation of the code in its final form was not published until 26 June 2018. The name of the ''Code'' is partly capitalized and partly not. The lower-case for "algae, fungi, and plants" indica ...
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Millettieae
The tribe Millettieae is one of the subdivisions of the plant family Fabaceae. The following genera are recognized by the USDA. In 2019, some genera USDA places in this tribe were moved to tribe Wisterieae; these are listed at the end. * ''Aganope'' Miq. * '' Antheroporum'' Gagnep. * '' Apurimacia'' Harms * '' Austrosteenisia'' R. Geesink * '' Bergeronia'' Micheli * '' Burkilliodendron'' Sastry * '' Chadsia'' Bojer * ''Craibia'' Harms & Dunn * '' Craspedolobium'' Harms * '' Dahlstedtia'' Malme * '' Dalbergiella'' Baker f. * '' Deguelia'' Aubl. * ''Derris'' Lour. * '' Dewevrea'' Micheli * '' Disynstemon'' R. Vig. * ''Fordia'' Hemsl. * '' Hesperothamnus'' Brandegee * ''Kunstleria'' Prain * '' Leptoderris'' Dunn * ''Lonchocarpus'' Kunth * '' Margaritolobium'' Harms * ''Millettia'' Wight & Arn. * '' Muellera'' L. f. * ''Mundulea'' (DC.) Benth. * '' Ostryocarpus'' Hook. f. * '' Paratephrosia'' Domin * ''Philenoptera'' Hochst. ''ex'' A.Rich. * '' Pi ...
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Trifolium
Clover or trefoil are common names for plants of the genus ''Trifolium'' (from Latin ''tres'' 'three' + ''folium'' 'leaf'), consisting of about 300 species of flowering plants in the legume or pea family Fabaceae originating in Europe. The genus has a cosmopolitan distribution with highest diversity in the temperate Northern Hemisphere, but many species also occur in South America and Africa, including at high altitudes on mountains in the tropics. They are small annual, biennial, or short-lived perennial herbaceous plants, typically growing up to 30 cm tall. The leaves are trifoliate (rarely quatrefoiled; see four-leaf clover), monofoil, bifoil, cinquefoil, hexafoil, septfoil, etcetera, with stipules adnate to the leaf-stalk, and heads or dense spikes of small red, purple, white, or yellow flowers; the small, few-seeded pods are enclosed in the calyx. Other closely related genera often called clovers include ''Melilotus'' (sweet clover) and '' Medicago'' (alfalfa or Calvary clover ...
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Swainsona
''Swainsona'' is a large genus of flowering plants native to Australasia. There are 85 species, all but one of which are endemic to Australia. A member of the family Fabaceae (legumes), it is most closely related to the New Zealand genera ''Montigena'' (scree pea), ''Clianthus'' (kakabeak), and ''Carmichaelia'' ( New Zealand broom). ''Swainsona'' is named after English botanist Isaac Swainson. A few species are known to produce swainsonine, a phytotoxin harmful to livestock (see Locoweed). In Australia, animals intoxicated with swainsonine are said to be pea struck. ;Selected species *'' Swainsona acuticarinata'' ( A.T.Lee) Joy Thomps. *'' Swainsona adenophylla'' J.M.Black *'' Swainsona affinis'' ( A.T.Lee) Joy Thomps. *'' Swainsona beasleyana'' F.Muell. *'' Swainsona behriana'' F.Muell. ex J.M.Black *'' Swainsona brachycarpa'' Benth. *'' Swainsona bracteata'' (Maiden & Betche) Joy Thomps. *'' Swainsona burkei'' F.Muell. ex Benth. *'' Swainsona burkittii'' F.Muell. ex Benth ...
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Oxytropis
''Oxytropis'' is a genus of plants in the legume family. It is one of three genera of plants known as locoweeds, and are notorious for being toxic to grazing animals. The other locoweed genus is the closely related ''Astragalus''. There are about 600 species, native to Eurasia and North America. Several species are native to the Arctic. These are hairy perennial plants which produce raceme inflorescences of pink, purple, white, or yellow flowers which are generally pea-like but have distinctive sharply beaked keels. The stems are leafless, the leaves being all basal. The plant produces legume pods containing the seeds. Selected species *'' Oxytropis arctica'' – Arctic locoweed *'' Oxytropis bellii'' *'' Oxytropis borealis'' – boreal locoweed *''Oxytropis campestris'' – field locoweed *'' Oxytropis deflexa'' – nodding locoweed *'' Oxytropis halleri'' – purple oxytropis *''Oxytropis jacquinii'' *'' Oxytropis kobukensis'' – Kobuk locoweed *'' Oxytropis lambertii'' – ...
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Medicago
''Medicago'' is a genus of flowering plants, commonly known as medick or burclover, in the legume family (Fabaceae). It contains at least 87 species and is distributed mainly around the Mediterranean basin. The best-known member of the genus is alfalfa (''M. sativa''), an important forage crop, and the genus name is based on the Latin name for that plant, , from el, μηδική (πόα) Median (grass). Most members of the genus are low, creeping herbs, resembling clover, but with burs (hence the common name). However, alfalfa grows to a height of 1 meter, and tree medick (''M. arborea'') is a shrub. Members of the genus are known to produce bioactive compounds such as medicarpin (a flavonoid) and medicagenic acid (a triterpenoid saponin). Chromosome numbers in ''Medicago'' range from 2''n'' = 14 to 48. The species ''Medicago truncatula'' is a model legume due to its relatively small stature, small genome (450–500 Mbp), short generation time (about 3 months), and ability to ...
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Hedysarum
''Hedysarum'' (sweetvetch) is a genus of the botanical family Fabaceae, consisting of about 200 species of annual or perennial herbs in Asia, Europe, North Africa, and North America. Description Species within genus ''Hedysarum'' may be herbaceous plants or deciduous shrubs. They have odd-pinnate leaves, with entire leaflets (no notches or indentations). These leaves resemble the leaves of sweet peas. The stipules may be free or connate, and stipels (secondary stipules) are absent. The inflorescences are peduncled racemes or heads. Bracts are small, with bracteoles below the calyx, and calyx teeth subequal. The petals may be pink, purplish, yellow, or whitish. Vexillum is longer than the wings, with an obtuse keel longer or rarely shorter than the wings. Stamens are diadelphous, 9+1, and anthers uniform. Ovary is 2-8-ovuled. Fruit is a lomentum, with segments that are glabrous, pubescent, bristly, or spiny, and break into single-seeded sections on ripening. Uses ''H ...
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Astragalus
''Astragalus'' is a large genus of over 3,000 species of herbs and small shrubs, belonging to the legume family Fabaceae and the subfamily Faboideae. It is the largest genus of plants in terms of described species. The genus is native to temperate regions of the Northern Hemisphere. Common names include milkvetch (most species), locoweed (in North America, some species) and goat's-thorn ( ''A. gummifer'', ''A. tragacantha''). Some pale-flowered vetches (''Vicia'' spp.) are similar in appearance, but they are more vine-like than ''Astragalus''. Description Most species in the genus have pinnately compound leaves. There are annual and perennial species. The flowers are formed in clusters in a raceme, each flower typical of the legume family, with three types of petals: banner, wings, and keel. The calyx is tubular or bell-shaped. Ecology ''Astragalus'' species are used as food plants by the larvae of some Lepidoptera species including many case-bearing moths of the genus ''Col ...
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Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope Carbon-13, 13C in the atmosphere was exceptionally low in comparison with the more common isotope Carbon-12, 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Popigai impact structure, Siberia and in what is now ...
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Systematic Biology
Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: cladograms, phylogenetic trees, phylogenies). Phylogenies have two components: branching order (showing group relationships) and branch length (showing amount of evolution). Phylogenetic trees of species and higher taxa are used to study the evolution of traits (e.g., anatomical or molecular characteristics) and the distribution of organisms (biogeography). Systematics, in other words, is used to understand the evolutionary history of life on Earth. The word systematics is derived from the Latin word '' systema,'' which means systematic arrangement of organisms. Carl Linnaeus used 'Systema Naturae' as the title of his book. Branches and applications In the study of biological systematics, researchers use the different branches to further understand the relationshi ...
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Systematic Botany
Plant taxonomy is the science that finds, identifies, describes, classifies, and names plants. It is one of the main branches of taxonomy (the science that finds, describes, classifies, and names living things). Plant taxonomy is closely allied to plant systematics, and there is no sharp boundary between the two. In practice, "plant systematics" involves relationships between plants and their evolution, especially at the higher levels, whereas "plant taxonomy" deals with the actual handling of plant specimens. The precise relationship between taxonomy and systematics, however, has changed along with the goals and methods employed. Plant taxonomy is well known for being turbulent, and traditionally not having any close agreement on circumscription and placement of taxa. See the list of systems of plant taxonomy. Background Classification systems serve the purpose of grouping organisms by characteristics common to each group. Plants are distinguished from animals by various trai ...
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