Struthiosaurini
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Struthiosaurini
Struthiosaurini (derived from ''Struthiosaurus'', "ostrich reptile") is a clade of nodosauridae, nodosaurid ankylosaurs from the Cretaceous of Europe and North America. The group is defined as the largest clade containing ''Struthiosaurus austriacus'', but not ''Nodosaurus textilis'' or ''Panoplosaurus mirus'', and was named in 2021 in paleontology, 2021 by Madzia and colleagues for the relatively stable group found in many previous analyses. Struthiosaurini includes not only the Late Cretaceous European ''Struthiosaurus'', but also the Early Cretaceous European ''Europelta'', the Late Cretaceous European ''Hungarosaurus'', and ''Stegopelta'' and ''Pawpawsaurus'' from the mid Cretaceous of North America. The approximately equivalent clade Struthiosaurinae, named in 1923 in paleontology, 1923 by Franz Nopcsa was previously used to include European nodosaurids, but was never formally named following the PhyloCode, so Madzia ''et al.'' named Struthiosaurini instead, as the group of taxa ...
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Struthiosaurus Austriacus
''Struthiosaurus'' (Latin ''struthio'' = ostrich + Greek language, Greek ''sauros'' = lizard) is a genus of Nodosauridae, nodosaurid dinosaurs, from the Late Cretaceous period (Santonian-Maastrichtian) of Austria, Romania, France and Hungary in Europe.''Struthiosaurus'' in The Dinosaur Encyclopaedia
at Dino Russ's Lair
It was a small dinosaur, measuring in length and weighing .


History of discovery

In 1859, geologist Eduard Suess at the ''Gute Hoffnung'' coal mine at Muthmannsdorf near Wiener Neustadt in Austria, discovered a dinosaur tooth on a stone pile. With the help of mine intendant Pawlowitsch it was attempted to find the source of the fossil material. The search proved fruitless at first but ultimately a t ...
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Struthiosaurus
''Struthiosaurus'' (Latin ''struthio'' = ostrich + Greek ''sauros'' = lizard) is a genus of nodosaurid dinosaurs, from the Late Cretaceous period (Santonian-Maastrichtian) of Austria, Romania, France and Hungary in Europe.''Struthiosaurus'' in The Dinosaur Encyclopaedia
at Dino Russ's Lair
It was a small dinosaur, measuring in length and weighing .


History of discovery

In 1859, geologist at the ''Gute Hoffnung'' coal mine at Muthmannsdorf near

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Europelta
''Europelta'' (meaning “Europe’s shield”) is a monospecific genus of nodosaurid dinosaur from Spain that lived during the Early Cretaceous (early Albian stage, ~113.0 Ma) in what is now the lower Escucha Formation of the Teruel Province. The type and only species, ''Europelta carbonensis'', is known from two associated partial skeletons, and represents the most complete ankylosaur known from Europe. ''Europelta'' was named in 2013 by James I. Kirkland and colleagues. ''Europelta'' has an estimated length of 5 metres (16 feet) and weight of 1.3 tonnes (2,866 lbs), making it the largest member of the clade Struthiosaurini.Paul, G.S., 2016, ''The Princeton Field Guide to Dinosaurs 2nd Edition'', Princeton University Press Discovery In 2011, the remains of two partial skeletons of an ankylosaur were discovered from the Fundación Conjunto Paleontológico of Teruel-Dinópolis locality AR-1 of the lower, located east of Ariño, in the northern Teruel Province in the Community o ...
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Pawpawsaurus
''Pawpawsaurus'', meaning "Pawpaw Lizard", is a nodosaurid ankylosaur from the Cretaceous (late Albian) of Tarrant County, Texas, discovered in May 1992. The only species yet assigned to this taxon, ''Pawpawsaurus campbelli,'' is based on a complete skull (lacking mandibles) from the marine Paw Paw Formation (Wachita Group). Discovery ''Pawpawsaurus'' was found in the Paw Paw Formation in Tarrant County, Texas, in May 1992, by Cameron Campbell. A complete skull is the only specimen, from which the binomial was named ''Pawpawsaurus campbelli''. The Paw Paw Formation has produced another nodosaurid, ''Texasetes pleurohalio'' (Coombs, 1995), which may prove to be a senior synonym of ''Pawpawsaurus''. This is the only nodosaurid known to have possessed the bony eyelids commonly found in ankylosaurids. The skull of ''Pawpawsaurus'' bears some notable similarities to that of ''Silvisaurus'', such as the presence of teeth in the premaxilla and the restriction of the osseous secondar ...
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Nodosaurus Textilis
''Nodosaurus'' (meaning "knobbed lizard") is a genus of herbivorous nodosaurid ankylosaurian dinosaur from the Late Cretaceous, the fossils of which are found exclusively in the Frontier Formation in Wyoming. Description ''Nodosaurus'' grew up to roughly long and it was an ornithischian dinosaur with bony dermal plates covering the top of its body, and it may have had spikes along its side as well. The dermal plates were arranged in bands along its body, with narrow bands over the ribs alternating with wider plates in between. These wider plates were covered in regularly arranged bony nodules, which give the animal its scientific name. In 2010 Paul estimated its length at 6 meters (20 ft) and its weight at 3.5 tonnes (3.85 short tons). It had four short legs, five-toed feet, a short neck, and a long, stiff, clubless tail. The head was narrow, with a pointed snout, powerful jaws, and small teeth. It perhaps ate soft plants, as it would have been unable to chew tough, fib ...
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Nodosaurus
''Nodosaurus'' (meaning "knobbed lizard") is a genus of herbivorous nodosaurid ankylosaurian dinosaur from the Late Cretaceous, the fossils of which are found exclusively in the Frontier Formation in Wyoming. Description ''Nodosaurus'' grew up to roughly long and it was an ornithischian dinosaur with bony dermal plates covering the top of its body, and it may have had spikes along its side as well. The dermal plates were arranged in bands along its body, with narrow bands over the ribs alternating with wider plates in between. These wider plates were covered in regularly arranged bony nodules, which give the animal its scientific name. In 2010 Paul estimated its length at 6 meters (20 ft) and its weight at 3.5 tonnes (3.85 short tons). It had four short legs, five-toed feet, a short neck, and a long, stiff, clubless tail. The head was narrow, with a pointed snout, powerful jaws, and small teeth. It perhaps ate soft plants, as it would have been unable to chew tough, fib ...
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Nodosaurinae
Nodosaurinae is a subfamily of nodosaurid ankylosaurs from the Cretaceous of Europe, North America, and South America. The group is defined as the largest clade containing '' Nodosaurus textilis'' but not ''Hylaeosaurus armatus'', '' Mymoorapelta maysi'', or ''Polacanthus foxii'', and was formally named in 2021 by Madzia and colleagues, who utilized the name of Othenio Abel in 1919, who created the term to unite ''Ankylosaurus'', ''Hierosaurus'' and ''Stegopelta''. The name has been significantly refined in content since Abel first used it to unite all quadrupedal, plate-armoured ornithischians, now including a significant number of taxa from the Early Cretaceous through Maastrichtian of Europe, North America, and Argentina. Previous informal definitions of the group described the clade as all taxa closer to ''Panoplosaurus'', or ''Panoplosaurus'' and ''Nodosaurus'', than to the early ankylosaurs ''Sarcolestes'', ''Hylaeosaurus'', ''Mymoorapelta'' or ''Polacanthus'', which was refle ...
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Acantholipan
''Acantholipan'' is a genus of herbivorous nodosaurid dinosaur from Mexico from the early Santonian age of the Late Cretaceous. It includes one species, ''Acantholipan gonzalezi.'' Discovery and naming In the north of Mexico, fragmentary fossils have been found of nodosaurids. A partial skeleton excavated at Los Primos near San Miguel in Coahuila, was described in 2011. When Rivera-Sylva and colleagues reported the discovery of this specimen, CPC 272, they initially considered it too fragmentary to name. Later it was judged that the remains were sufficiently distinct to be given a binomial name. In 2018, the type species ''Acantholipan gonzalezi'' was named by Héctor Eduardo Rivera-Sylva, Eberhard Frey, Wolfgang Stinnesbeck, Gerardo Carbot-Chanona, Iván Erick Sanchez-Uribe and José Rubén Guzmán-Gutiárrez. The generic name combines a Greek ''akanthos'', "spine", with ''lipan'', the usual Spanish designation of the '' Lépai-Ndé'', the "Gray People", a tribe of the Ap ...
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Peloroplites
''Peloroplites'' (meaning “monstrous heavy one”) is a monospecific genus of nodosauridae, nodosaurid dinosaur from Utah that lived during the Late Cretaceous (Cenomanian to lower Turonian stage, 98.2 to 93 Ma) in what is now the Mussentuchit Member of the Cedar Mountain Formation. The type and only species, ''Peloroplites cedrimontanus'', is known from a partial skull and postcrania, postcranial skeleton. It was named in 2008 by Kenneth Carpenter and colleagues. ''Peloroplites'' was 6 metres (20 feet) long and weighed 2 tonnes (4,410 lbs), making it one of the largest known nodosaurids, and came from a time when ankylosauridae, ankylosaurids and nodosaurids were attaining large sizes.Paul, G.S., 2016, ''The Princeton Field Guide to Dinosaurs 2nd Edition'', Princeton University Press Discovery and Naming In 2001, a skeleton of a large nodosauridae, nodosaurid from the Cedar Mountain Formation in Emery County, Utah, Emery County, Utah was mentioned by Burge and Bird in a publi ...
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Panoplosaurus Mirus
''Panoplosaurus'' is a genus of armoured dinosaur from the Late Cretaceous of Alberta, Canada. Few specimens of the genus are known, all from the middle Campanian of the Dinosaur Park Formation, roughly 76 to 75 million years ago. It was first discovered in 1917, and named in 1919 by Lawrence Lambe, named for its extensive armour, meaning "well-armoured lizard". ''Panoplosaurus'' has at times been considered the proper name for material otherwise referred to as ''Edmontonia'', complicating its phylogenetic and ecological interpretations, at one point being considered to have existed across Alberta, New Mexico and Texas, with specimens in institutions from Canada and the United States. The skull and skeleton of ''Panoplosaurus'' are similar to its relatives, but have a few significant differences, such as the lumpy form of the skull osteoderms, a completely fused shoulder blade, and regularly shaped plates on its neck and body lacking prominent spines. It was a quadrupedal animal, ...
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Panoplosaurini
Panoplosaurini (derived from ''Panoplosaurus'', "all shield reptile") is a clade of nodosaurid ankylosaurs from the Cretaceous of North America and South America. The group is defined as the largest clade containing ''Panoplosaurus mirus'', but not '' Nodosaurus textilis'' or '' Struthiosaurus austriacus'', and was named in 2021 by Madzia and colleagues for the group found in many previous analyses, both morphological and phylogenetic. Panoplosaurini includes not only the Late Cretaceous ''Panoplosaurus'', ''Denversaurus'' and ''Edmontonia'', but also the mid Cretaceous ''Animantarx'' and ''Texasetes'', as well as ''Patagopelta''. However, in the study describing it, its authors only placed it as a nodosaurine outside Panoplosaurini. The approximately equivalent clade Panoplosaurinae, named in 1929 by Franz Nopcsa, but was not significantly used until Robert Bakker reused the name in 1988, alongside the new clades Edmontoniinae and Edmontoniidae, which were considered to unite ''Pan ...
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Nodosauridae
Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, were heavily armored dinosaurs adorned with rows of bony armor nodules and spines (osteoderms), which were covered in keratin sheaths. All nodosaurids, like other ankylosaurians, were medium-sized to large, heavily built, quadrupedal, herbivorous dinosaurs, possessing small, leaf-shaped teeth. Unlike ankylosaurids, nodosaurids lacked mace-like tail clubs, instead having flexible tail tips. Many nodosaurids had spikes projecting outward from their shoulders. One particularly well-preserved nodosaurid "mummy", known as the Suncor nodosaur (''Borealopelta markmitchelli''), preserved a nearly complete set of armor in life position, as well as the keratin covering and mineralized remains of the underlying skin, which indicate reddish dorsal pigmen ...
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