Struthiosaurini
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Struthiosaurini (derived from ''
Struthiosaurus ''Struthiosaurus'' (Latin ''struthio'' = ostrich + Greek ''sauros'' = lizard) is a genus of nodosaurid dinosaurs, from the Late Cretaceous period (Santonian-Maastrichtian) of Austria, Romania, France and Hungary in Europe.
'', "ostrich reptile") is a clade of
nodosaurid Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, wer ...
ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
s from the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
of
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
and
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. The group is defined as the largest clade containing ''
Struthiosaurus austriacus ''Struthiosaurus'' (Latin ''struthio'' = ostrich + Greek language, Greek ''sauros'' = lizard) is a genus of Nodosauridae, nodosaurid dinosaurs, from the Late Cretaceous period (Santonian-Maastrichtian) of Austria, Romania, France and Hungary in E ...
'', but not ''
Nodosaurus textilis ''Nodosaurus'' (meaning "knobbed lizard") is a genus of herbivorous nodosaurid ankylosaurian dinosaur from the Late Cretaceous, the fossils of which are found exclusively in the Frontier Formation in Wyoming. Description ''Nodosaurus'' grew ...
'' or ''
Panoplosaurus mirus ''Panoplosaurus'' is a genus of armoured dinosaur from the Late Cretaceous of Alberta, Canada. Few specimens of the genus are known, all from the middle Campanian of the Dinosaur Park Formation, roughly 76 to 75 million years ago. It was first d ...
'', and was named in
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by Madzia and colleagues for the relatively stable group found in many previous analyses. Struthiosaurini includes not only the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
European ''Struthiosaurus'', but also the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
European ''
Europelta ''Europelta'' (meaning “Europe’s shield”) is a monospecific genus of nodosaurid dinosaur from Spain that lived during the Early Cretaceous (early Albian stage, ~113.0 Ma) in what is now the lower Escucha Formation of the Teruel Province. T ...
'', the Late Cretaceous European ''
Hungarosaurus ''Hungarosaurus tormai'' is a herbivorous nodosaurid ankylosaur from the Upper Cretaceous (Santonian) Csehbánya Formation of the Bakony Mountains of western Hungary. It is the most completely known ankylosaur from the Cretaceous of Europe. Di ...
'', and ''
Stegopelta ''Stegopelta'' (meaning "roofed shield") is a genus of struthiosaurin nodosaurid dinosaur. It is based on a partial skeleton from the latest Albian- earliest Cenomanian-age Lower and Upper Cretaceous Belle Fourche Member of the Frontier Format ...
'' and ''
Pawpawsaurus ''Pawpawsaurus'', meaning "Pawpaw Lizard", is a nodosaurid ankylosaur from the Cretaceous (late Albian) of Tarrant County, Texas, discovered in May 1992. The only species yet assigned to this taxon, ''Pawpawsaurus campbelli,'' is based on a comp ...
'' from the mid Cretaceous of North America. The approximately equivalent clade Struthiosaurinae, named in
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by
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was previously used to include European nodosaurids, but was never formally named following the
PhyloCode The ''International Code of Phylogenetic Nomenclature'', known as the ''PhyloCode'' for short, is a formal set of rules governing phylogenetic nomenclature. Its current version is specifically designed to regulate the naming of clades, leaving the ...
, so Madzia ''et al.'' named Struthiosaurini instead, as the group of taxa fell within the clade
Nodosaurinae Nodosaurinae is a subfamily of nodosaurid ankylosaurs from the Cretaceous of Europe, North America, and South America. The group is defined as the largest clade containing '' Nodosaurus textilis'' but not ''Hylaeosaurus armatus'', '' Mymoorapelta ...
, and having the same ''-inae'' suffix on both parent and child taxon could be confusing in future. The
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phylogenetic analysis of Rivera-Sylva and colleagues was used as the primary reference for Struthiosaurini by Madzia ''et al.'', in addition to the supplemental analyses of Arbour ''et al.'' (2016), Brown ''et al.'' (2017), and Zheng ''et al.'' (2018).


References

{{Thyreophora Nodosaurids Cretaceous dinosaurs