Simoliophiidae
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Simoliophiidae
Simoliophiidae is an extinct family of limbed Tethyan marine snakes of the order Squamata Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest order of reptiles, comprising lizards, snakes, and amphisbaenians (worm lizards), which are collectively known as squamates or scaled reptiles. With over 10,900 species, .... The name Pachyophiidae has also been used for this group, but Simoliophiidae has priority. References Prehistoric snakes Marine reptiles Alethinophidia Prehistoric reptile families {{snake-stub ...
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Pachyophis
''Pachyophis'' is an extinct genus of Simoliophiidae snakes that were extant during the Cenomanian stage of the Late Cretaceous period. More specifically, it was found to be from the Cenomanian Age about 93.9-100.5 million years ago in the suburb area of Bileca, Herzegovina. ''Pachyophis'' belongs to the family Simoliophiidae (hind-limbed snakes) in the clade Ophidia. The family is characterized by the presence of hindlimbs and pelvic girdles, as seen in ''Pachyrhachis'' (one of the first discovered hind-limbed snakes). Included in the most basal of snake relatives, ''Pachyophis'' in contrast does not have evidence of any form of limb preservation. Despite this, they are considered under Pachyophiidae due to their other morphological similarities along with the genera '' Mesophis'' and '' Simoliophis''. Only one species has been discovered, which is known as ''Pachyophis woodwardi''. Discovery There is a single species discovered of ''Pachyophis'', found in 1923 by F. Nopsc ...
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Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding age. Both age and stage bear the same name. As a unit of geologic time measure, the Cenomanian Age spans the time between 100.5 and 93.9 million years ago (Mya). In the geologic timescale, it is preceded by the Albian and is followed by the Turonian. The Upper Cenomanian starts around at 95 Mya. The Cenomanian is coeval with the Woodbinian of the regional timescale of the Gulf of Mexico and the early part of the Eaglefordian of the regional timescale of the East Coast of the United States. At the end of the Cenomanian, an anoxic event took place, called the Cenomanian-Turonian boundary event or the "Bonarelli event", that is associated with a minor extinction event for marine spec ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Eupodophis
''Eupodophis'' is an extinct genus of snake from the Late Cretaceous period. It has two small hind legs and is considered a transitional form between Cretaceous lizards and limbless snakes. The feature, described as vestigial, was most likely useless to ''Eupodophis''. The type species ''Eupodophis descouensi'' was named in 2000 and resides now in the paleontology section of the Mim Museum in Beirut, Lebanon. The specific name is dedicated to the French naturalist Didier Descouens. The fossilized specimen from which the description of the type species was based was 85 cm (33.5 in) long and is approximately 92 million years old. It was found in Cenomanian-age limestone near the al-Nammoura village in Lebanon belonging to the Sannine Formation. Description ''Eupodophis'' was a marine snake that lived in the Mediterranean Tethys Ocean. It had a laterally compressed body and short, paddle-like tail. The vertebrae and ribs of ''Eupodophis'' are pachyostotic, or thickened, ...
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Haasiophis
''Haasiophis'', consisting of the sole species ''Haasiophis terrasanctus'', is an extinct genus of snakes with hind limbs. It is one of three genera of Cenomanian snakes known to have possessed hindlimbs. Etymology The genus was named in honor of the late paleontologist Georg Haas, who first began work in the fossils of Ein Yabrud and started the description of the genus before he died, plus the Greek "ophis", for snake . The species names is from the Latin "terrasanctus" meaning "holy land". Specimen material ''H. terrasanctus'' is known from a single fossil discovered at Ein Yabrud in the Judean hills, near Ramallah, 20 km north of Jerusalem, in the central West Bank. This site also produced the type and only specimen for the sister genus ''Pachyrhachis''. et al. 2003 "The Anatomy and Relationships of Haasiophis terrasanctus, a Fossil Snake with Well-Developed Hind Limbs from the Mid-Cretaceous of the Middle East" ''Journal of Paleontology'' 77(3):536-558 This site i ...
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Pachyrhachis
''Pachyrhachis'' (from el, παχύς , 'thick' and el, ῥάχῐς , 'spine') is an extinct genus of snake with well developed hind legs known from fossils discovered in Ein Yabrud, near Ramallah, in the central West Bank. It is a relatively small snake, measuring more than long at maximum. ''Pachyrhachis'' appears to have been an ancient marine snake; the fossils occur in a marine limestone deposit, and the thickened bone of the ribs and vertebrae would have functioned as ballast to decrease the buoyancy of the animal, allowing it to dive beneath the ancient Cretaceous seas that it once inhabited. ''Pachyrhachis'' is one of three genera of Cenomanian snakes with hindlimbs. Although many modern pythons and boas still retain remnants of legs, in the form of small spurs, the tiny legs of Pachyrhachis included a hip, knee, and ankle joint. ''Pachyrhachis'' was originally described by Haas (1979, 1980) who noted it had a puzzling melange of snake and lizard features; its status a ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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Tethys Ocean
The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents of Gondwana and Laurasia, before the opening of the Indian and Atlantic oceans during the Cretaceous Period. It was preceded by the Paleo-Tethys Ocean, which lasted between the Cambrian and the Early Triassic, while the Neotethys formed during the Late Triassic and lasted until the early Eocene (about 50 million years ago) when it completely closed. A portion known as the Paratethys formed during the Late Jurassic, was isolated during the Oligocene (34 million years ago) and lasted up to the Pliocene (about 5 million years ago), when it largely dried out. Many major seas and lakes of Europe and Western Asia, including the Mediterranean Sea, the Black Sea, the Caspian Sea, and the Aral Sea are thought to be remnants of the Paratethys. Ety ...
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Marine Reptile
Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. The earliest marine reptile mesosaurus (not to be confused with mosasaurus), arose in the Permian period during the Paleozoic era. During the Mesozoic era, many groups of reptiles became adapted to life in the seas, including such familiar clades as the ichthyosaurs, plesiosaurs (these two orders were once thought united in the group "Enaliosauria", a classification now cladistically obsolete), mosasaurs, nothosaurs, placodonts, sea turtles, thalattosaurs and thalattosuchians. Most marine reptile groups became extinct at the end of the Cretaceous period, but some still existed during the Cenozoic, most importantly the sea turtles. Other Cenozoic marine reptiles included the bothremydids, palaeophiid snakes, a few choristoderes such as ''Simoedosaurus'' and dyrosaurid crocodylomorphs. Various types of marine gavialid crocodilians remained widespread as ...
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Squamata
Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest order of reptiles, comprising lizards, snakes, and amphisbaenians (worm lizards), which are collectively known as squamates or scaled reptiles. With over 10,900 species, it is also the second-largest order of extant (living) vertebrates, after the perciform fish. Members of the order are distinguished by their skins, which bear horny scales or shields, and must periodically engage in molting. They also possess movable quadrate bones, making possible movement of the upper jaw relative to the neurocranium. This is particularly visible in snakes, which are able to open their mouths very wide to accommodate comparatively large prey. Squamata is the most variably sized order of reptiles, ranging from the dwarf gecko (''Sphaerodactylus ariasae'') to the Reticulated python (''Malayopython reticulatus'') and the now-extinct mosasaurs, which reached lengths over . Among other reptiles, squamates are most close ...
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