Scleroglossa
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Scleroglossa
Scleroglossa is a group of lizards that includes geckos, autarchoglossans (scincomorphs, anguimorphs, and varanoids), and amphisbaenians. Scleroglossa is supported by phylogenetic analyses that use morphological features (visible anatomical features). According to most morphological analyses, Scleroglossa is the sister group of the clade Iguania, which includes iguanas, chameleons, agamids, and New World lizards. Together, Scleroglossa and Iguania make up the crown group Squamata, the smallest evolutionary grouping to include all living lizards and snakes. The name Scleroglossa is derived from the Greek, ''skleros'', meaning "hard" and ''glossa'', meaning "tongue". The split between Scleroglossa and Iguania can be based on features of the tongue; iguanians have a muscular tongue and use lingual prehension to capture food, whereas scleroglossans have hard tongues and use teeth-and-jaw prehension to capture food, freeing the tongue for chemosensory activity. Recent phylogenetic ana ...
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Scincogekkonomorpha
Scincogekkonomorpha is a clade (evolutionary grouping) of lizards that includes scleroglossans and all lizards more closely related to scleroglossans than to iguanians. These "stem" scleroglossans include extinct lizards from the Late Jurassic and Early Cretaceous such as ''Bavarisaurus'', ''Eichstaettisaurus'', '' Liushusaurus'', and '' Scandensia''. Scincogekkonomorpha was named in 1961 and is now occasionally used as a stem-based taxon in contrast to the node-based taxon Scleroglossa. According to phylogenies based on morphological characteristics, Scincogekkonomorpha is the sister taxon of Iguania and together they make up crown group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. ... Squamata, the smallest clade including all living snakes and lizards. References Tetra ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Amphisbaenian
Amphisbaenia (called amphisbaenians or worm lizards) is a group of usually legless squamates, comprising over 200 extant species. Amphisbaenians are characterized by their long bodies, the reduction or loss of the limbs, and rudimentary eyes. As many species have a pink body and scales arranged in rings, they have a superficial resemblance to earthworms. While the genus '' Bipes'' retains forelimbs, all other genera are limbless. Although superficially similar to the snakes and blind lizards, recent phylogenetic studies suggest that they are most closely related to wall lizards of the family Lacertidae. Amphisbaenians are widely distributed, occurring in North America, Europe, Africa, South America, Western Asia and the Caribbean. Most species are less than long. Description 200px, left, Close-up of the head of'' Rhineura'' Despite a superficial resemblance to some primitive snakes, amphisbaenians have many unique features that distinguish them from other reptiles. Internal ...
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Node-based Taxon
Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which can be a specimen or a taxon of lower rank, and a description in words. Phylogenetic nomenclature is currently regulated by the '' International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature ties names to clades, groups consisting of an ancestor and all its descendants. These groups can equivalently be called monophyletic. There are slightly different ways of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the named taxon. Listing all these organisms (i.e. providing a full circumscription) requires the full phylogenetic tree to be known. In practice, ther ...
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Stem-based Taxon
Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which can be a specimen or a taxon of lower rank, and a description in words. Phylogenetic nomenclature is currently regulated by the '' International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature ties names to clades, groups consisting of an ancestor and all its descendants. These groups can equivalently be called monophyletic. There are slightly different ways of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the named taxon. Listing all these organisms (i.e. providing a full circumscription) requires the full phylogenetic tree to be known. In practice, the ...
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Snakes
Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads (cranial kinesis). To accommodate their narrow bodies, snakes' paired organs (such as kidneys) appear one in front of the other instead of side by side, and most have only one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca. Lizards have evolved elongate bodies without limbs or with greatly reduced limbs about twenty-five times independently via convergent evolution, leading to many lineages of legless lizards. These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, although this rule is not universal (see Amphisbaenia, D ...
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Crown Group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown gr ...
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Agamid
Agamidae is a family of over 300 species of iguanian lizards indigenous to Africa, Asia, Australia, and a few in Southern Europe. Many species are commonly called dragons or dragon lizards. Overview Phylogenetically, they may be sister to the Iguanidae, and have a similar appearance. Agamids usually have well-developed, strong legs. Their tails cannot be shed and regenerated like those of geckos (and several other families such as skinks), though a certain amount of regeneration is observed in some. Many agamid species are capable of limited change of their colours to regulate their body temperature. In some species, males are more brightly coloured than females, and colours play a part in signaling and reproductive behaviours. Although agamids generally inhabit warm environments, ranging from hot deserts to tropical rainforests, at least one species, the mountain dragon, is found in cooler regions. They are particularly diverse in Australia. This group of lizards includes som ...
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Chameleon
Chameleons or chamaeleons (family Chamaeleonidae) are a distinctive and highly specialized clade of Old World lizards with 202 species described as of June 2015. The members of this family are best known for their distinct range of colors, being capable of shifting to different hues and degrees of brightness. The large number of species in the family exhibit considerable variability in their capacity to change color. For some, it is more of a shift of brightness (shades of brown); for others, a plethora of color-combinations (reds, yellows, greens, blues) can be seen. Chameleons are distinguished by their zygodactylous feet, their prehensile tail, their laterally compressed bodies, their head casques, their projectile tongues, their swaying gait, and crests or horns on their brow and snout. Chameleons' eyes are independently mobile, and because of this there are two separate, individual images that the brain is analyzing of the chameleon’s environment. When hunting prey, they ...
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Iguana
''Iguana'' (, ) is a genus of herbivorous lizards that are native to tropical areas of Mexico, Central America, South America, and the Caribbean. The genus was first described in 1768 by Austrian naturalist Josephus Nicolaus Laurenti in his book ''Specimen Medicum, Exhibens Synopsin Reptilium Emendatam cum Experimentis circa Venena''. Two species are placed in the genus, the green iguana, which is widespread throughout its range and a popular pet, and the Lesser Antillean iguana, which is native to the Lesser Antilles. Genetic analysis indicates that the green iguana may comprise a complex of multiple species, some of which have been recently described, but the Reptile Database considers all of these as subspecies of the green iguana. The word "iguana" is derived from the original Taino name for the species, ''iwana''. In addition to the two species in the genus ''Iguana'', several other related genera in the same family have common names of the species including the word "ig ...
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Iguania
Iguania is an infraorder of squamate reptiles that includes iguanas, chameleons, agamids, and New World lizards like anoles and phrynosomatids. Using morphological features as a guide to evolutionary relationships, the Iguania are believed to form the sister group to the remainder of the Squamata, which comprise nearly 11,000 named species, roughly 2000 of which are iguanians. However, molecular information has placed Iguania well within the Squamata as sister taxa to the Anguimorpha and closely related to snakes. The order has been under debate and revisions after being classified by Charles Lewis Camp in 1923 due to difficulties finding adequate synapomorphic morphological characteristics. Most Iguanias are arboreal but there are several terrestrial groups. They usually have primitive fleshy, non-prehensile tongues, although the tongue is highly modified in chameleons. The group has a fossil record that extends back to the Early Jurassic (the oldest known member is '' B ...
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Sister Group
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomic ...
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