Surusicaris
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Surusicaris
''Surusicaris'' is an extinct genus of bivalved arthropod, known from the Cambrian Burgess Shale of British Columbia, Canada. It is considered to be closely related to ''Isoxys,'' and like it has spined grasping frontal appendages. Description The only known specimen (ROM 62976), has a roughly semicircular bivalved carapace, which is approximately long and tall. The head has a pair of rounded eyes, and a pair of upward-curling segmented frontal appendages with at least five segments, the second, third and fourth of which bear a spine, with three long spines and at least one short spine on the fifth and terminal segment. The head also has three pairs of other limbs, which are stubby, uniramous and unsclerotised, with an annulated appearance, which are tipped with bifid claws. Along the trunk are 12 pairs of biramous limbs, which taper in size posteriorly. Their morphology is simple, the morphology of the endopod is similar to those of the anterior three uniramous head limbs, whi ...
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Surusicaris Restoration
''Surusicaris'' is an extinct genus of bivalved arthropod, known from the Cambrian Burgess Shale of British Columbia, Canada. It is considered to be closely related to ''Isoxys,'' and like it has spined grasping frontal appendages. Description The only known specimen (ROM 62976), has a roughly semicircular bivalved carapace, which is approximately long and tall. The head has a pair of rounded eyes, and a pair of upward-curling segmented frontal appendages with at least five segments, the second, third and fourth of which bear a spine, with three long spines and at least one short spine on the fifth and terminal segment. The head also has three pairs of other limbs, which are stubby, uniramous and unsclerotised, with an annulated appearance, which are tipped with bifid claws. Along the trunk are 12 pairs of biramous limbs, which taper in size posteriorly. Their morphology is simple, the morphology of the endopod is similar to those of the anterior three uniramous head limbs, whi ...
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Isoxys
''Isoxys'' (meaning "equal surfaces") is a genus of extinct bivalved Cambrian arthropod; the various species of which are thought to have been freely swimming predators. It had a pair of large spherical eyes (which are the most commonly preserved feature of the soft-bodied anatomy), and two large frontal appendages used to grasp prey. Description Species of ''Isoxys'' have roughly semicircular bivalved carapaces, which vary in morphology between species. The front and rear edges of the carapaces bear foward and posterior facing spines, respectively which in some species are greatly elongated''.'' The carapaces of ''Isoxys'' are typically in length, excluding the spines, though some species are known to reach over . In long-spined species when including spine length, some specimens exceed . The opening angle of the carapace was close to vertical, giving it a narrow profile when viewed from above.'''' The head had a pair of large spherical stalked eyes, as well as a pair of upwa ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Burgess Shale
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. The rock unit is a black shale and crops out at a number of localities near the town of Field in Yoho National Park Yoho National Park ( ) is a National Parks of Canada, national park of Canada. It is located within the Canadian Rockies, Rocky Mountains along the western slope of the Continental Divide of the Americas in southeastern British Columbia, bordered ... and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42 km to the south. History and significance The Burgess Shale was discovered by palaeontologist Charles Doolittle Walcott, Charles Walcott on 30 August 1909, towards the end of the season's fieldwork. He returned in 1910 with his sons, daughter, and wif ...
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Uniramous
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, plural ''coxae''), ''trochanter'', ''femur'' (plural ''femora''), ''tibia'' (plural ''tibiae''), ''tarsus'' (plural ''tarsi''), ''ischium'' (plural ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (plural ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments artic ...
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Biramous
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, plural ''coxae''), ''trochanter'', ''femur'' (plural ''femora''), ''tibia'' (plural ''tibiae''), ''tarsus'' (plural ''tarsi''), ''ischium'' (plural ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (plural ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments artic ...
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Endopod
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, plural ''coxae''), ''trochanter'', ''femur'' (plural ''femora''), ''tibia'' (plural ''tibiae''), ''tarsus'' (plural ''tarsi''), ''ischium'' (plural ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (plural ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments artic ...
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Exopod
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, plural ''coxae''), ''trochanter'', ''femur'' (plural ''femora''), ''tibia'' (plural ''tibiae''), ''tarsus'' (plural ''tarsi''), ''ischium'' (plural ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (plural ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments artic ...
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Seta
In biology, setae (singular seta ; from the Latin word for "bristle") are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Annelid setae are stiff bristles present on the body. They help, for example, earthworms to attach to the surface and prevent backsliding during peristaltic motion. These hairs make it difficult to pull a worm straight from the ground. Setae in oligochaetes (a group including earthworms) are largely composed of chitin. They are classified according to the limb to which they are attached; for instance, notosetae are attached to notopodia; neurosetae to neuropodia. Crustaceans have mechano- and chemosensory setae. Setae are especially present on the mouthparts of crustaceans and can also be found on grooming limbs. In some cases, setae are modified into scale like structures. Setae on the legs of krill and other small crustaceans help them to gather phytoplankton. It captures them and allows th ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Burgess Shale Fossils
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's book '' Wonderful Life'' describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested. The fossil beds are in a series of shale layers, averaging and totalling about in thickness. These layers were deposited against the face of a high undersea limestone cliff. All these features were later raised up above current sea level during the cr ...
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