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Stephanorhinus
''Stephanorhinus'' is an extinct genus of two-horned rhinoceros native to Eurasia and North Africa that lived during the Pliocene to Late Pleistocene. Species of ''Stephanorhinus'' were the predominant and often only species of rhinoceros in much of temperate Eurasia, especially Europe, for most of the Pleistocene. Two species of ''Stephanorhinus'' – Merck's rhinoceros (''S. kirchbergensis'') and the narrow-nosed rhinoceros (''S. hemitoechus'') – persisted into the last glacial period. Etymology The first part of the name, ''Stephano-'', honours Stephen I, the first king of Hungary. (The genus name was coined by Kretzoi, a Hungarian.) The second part is from (Greek for "nose"), a typical suffix of rhinoceros genus names. Taxonomy The taxonomic history of ''Stephanorhinus'' is long and convoluted, as many species are known by numerous synonyms and different genera – typically ''Rhinoceros'' and ''Dicerorhinus'' – for the 19th and most of the early 20th century. Th ...
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Stephanorhinus Etruscus Skull
''Stephanorhinus'' is an extinct genus of two-horned rhinoceros native to Eurasia and North Africa that lived during the Pliocene to Late Pleistocene. Species of ''Stephanorhinus'' were the predominant and often only species of rhinoceros in much of temperate Eurasia, especially Europe, for most of the Pleistocene. Two species of ''Stephanorhinus'' – Merck's rhinoceros (''S. kirchbergensis'') and the narrow-nosed rhinoceros (''S. hemitoechus'') – persisted into the last glacial period. Etymology The first part of the name, ''Stephano-'', honours Stephen I, the first king of Hungary. (The genus name was coined by Kretzoi, a Hungarian.) The second part is from (Greek for "nose"), a typical suffix of rhinoceros genus names. Taxonomy The taxonomic history of ''Stephanorhinus'' is long and convoluted, as many species are known by numerous synonyms and different genera – typically ''Rhinoceros'' and ''Dicerorhinus'' – for the 19th and most of the early 20th century. Th ...
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Stephanorhinus Jeanvireti Skull
''Stephanorhinus'' is an extinct genus of two-horned rhinoceros native to Eurasia and North Africa that lived during the Pliocene to Late Pleistocene. Species of ''Stephanorhinus'' were the predominant and often only species of rhinoceros in much of temperate Eurasia, especially Europe, for most of the Pleistocene. Two species of ''Stephanorhinus'' – Merck's rhinoceros (''S. kirchbergensis'') and the narrow-nosed rhinoceros (''S. hemitoechus'') – persisted into the last glacial period. Etymology The first part of the name, ''Stephano-'', honours Stephen I, the first king of Hungary. (The genus name was coined by Kretzoi, a Hungarian.) The second part is from (Greek for "nose"), a typical suffix of rhinoceros genus names. Taxonomy The taxonomic history of ''Stephanorhinus'' is long and convoluted, as many species are known by numerous synonyms and different genera – typically ''Rhinoceros'' and ''Dicerorhinus'' – for the 19th and most of the early 20th century. Th ...
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Merck's Rhinoceros
''Stephanorhinus kirchbergensis'', also known as Merck's rhinoceros or the forest rhinoceros, is an extinct species of rhino known from the Middle to Late Pleistocene of Eurasia. One of the last members of the genus ''Stephanorhinus'', it is considered to be a typical component of the interglacial ''Palaeoloxodon'' large faunal assemblage in Europe. Among extant species of rhinoceroses it is most closely related to the Sumatran rhinoceros, while the well known woolly rhinoceros was another close relative. In the western part of its range, it was sympatric with ''Stephanorhinus hemitoechus''. Etymology and taxonomy The first part of the genus name is derived from that of King Stephen I of Hungary, and the second part from 'rhinos' (ρινος, meaning "nose"), as with ''Dicerorhinus.'' The species name was given by Georg Friedrich von Jäger in 1839 for Kirchberg an der Jagst in Baden-Württemberg, Germany where the type specimens had been found. It is often known in English ( ...
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Narrow-nosed Rhinoceros
The narrow-nosed rhinoceros (''Stephanorhinus hemitoechus'') is an extinct species of rhinoceros that lived in western Eurasia and North Africa during the Pleistocene. It first appeared in Europe some 600,000 years ago, and survived there until as recently as 40,000 years Before Present Description The narrow-nosed rhinoceros was a large animal, reaching a shoulder height of as much as . It can be distinguished from other species of ''Stephanorhinus'' by its very long and low skull. Its nasals are relatively low, and its horn base poorly developed. Teeth are forward shifting. The narrow-nosed rhinoceros probably favored temperate open areas rich in low-growing vegetation. It displayed many similarities to its better known extinct relative, the woolly rhinoceros. However, it was probably not a true grazer, but a mixed feeder, eating low-growing vegetation in open habitats. In Apulia in southern Italy, remains of narrow-nosed rhinoceros from the middle Late Pleistocene were fo ...
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Sumatran Rhinoceros
The Sumatran rhinoceros (''Dicerorhinus sumatrensis''), also known as the Sumatran rhino, hairy rhinoceros or Asian two-horned rhinoceros, is a rare member of the family Rhinocerotidae and one of five extant species of rhinoceros. It is the only extant species of the genus ''Dicerorhinus''. It is the smallest rhinoceros, although it is still a large mammal; it stands high at the shoulder, with a head-and-body length of and a tail of . The weight is reported to range from , averaging . Like both African species, it has two horns; the larger is the nasal horn, typically , while the other horn is typically a stub. A coat of reddish-brown hair covers most of the Sumatran rhino's body. The Sumatran rhinoceros once inhabited rainforests, swamps and cloud forests in India, Bhutan, Bangladesh, Myanmar, Laos, Thailand, Malaysia, Indonesia and southwestern China, particularly in Sichuan. It is now critically endangered, with only five substantial populations in the wild: four in Suma ...
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Rhinoceros
A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea.) Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh at least one tonne in adulthood. They have a herbivorous diet, small brains (400–600 g) for mammals of their size, one or two horns, and a thick (1.5–5 cm), protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their lips to pl ...
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Coelodonta
''Coelodonta'' (, from the Greek κοιλία, ''koilía'' and οδούς, ''odoús'', "hollow tooth", in reference to the deep grooves of their molars) is an extinct genus of rhinoceros that lived in Eurasia between 3.7 million years to 10,000 years before the present, in the Pliocene and the Pleistocene epochs. It is best known from the type species, the woolly rhinoceros (''Coelodonta antiquitatis''), which ranged throughout northern Eurasia during the Pleistocene. The earliest known species, ''Coelodonta thibetana,'' lived in Tibet during the Pliocene, with the genus spreading to the rest of Eurasia during the Pleistocene. Species Species recognised as members of ''Coelodonta'', according to Deng ''et al''. (2011), include: * ''Coelodonta thibetana'' (Deng et al. 2011): The most primitive species of the genus, inhabited the Tibetan Plateau during the Pliocene. * ''Coelodonta nihowanensis'' (Chow, 1978): A primitive species from northern China, it lived in the earliest Ple ...
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Woolly Rhinoceros
The woolly rhinoceros (''Coelodonta antiquitatis'') is an extinct species of rhinoceros that was common throughout Europe and Asia during the Pleistocene epoch and survived until the end of the last glacial period. The woolly rhinoceros was a member of the Pleistocene megafauna. The woolly rhinoceros was covered with long, thick hair that allowed it to survive in the extremely cold, harsh mammoth steppe. It had a massive hump reaching from its shoulder and fed mainly on herbaceous plants that grew in the steppe. Mummified carcasses preserved in permafrost and many bone remains of woolly rhinoceroses have been found. Images of woolly rhinoceroses are found among cave paintings in Europe and Asia. Taxonomy Woolly rhinoceros remains have been known long before the species was described, and were the basis for some mythical creatures. Native peoples of Siberia believed their horns were the claws of giant birds. A rhinoceros skull was found in Kl ...
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Dicerorhinus
''Dicerorhinus'' (Greek: "two" (dio), "horn" (keratos), "nose" (rhinos)) is a genus of the family Rhinocerotidae, consisting of a single extant species, the two-horned Sumatran rhinoceros (''D. sumatrensis''), and several extinct species. The genus likely originated in the Mid to Late Pliocene of Northern Indochina and South China. Many species attributed to the genus probably actually belong in ''Stephanorhinus.'' Taxonomy Historically, ''Dicerorhinus'' was a wastebasket taxon. Revisions by several authors over the years have removed many species: Transferred to ''Stephanorhinus'' *''Dicerorhinus merckii'' *''Dicerorhinus hemitoechus'' *''Dicerorhinus etruscus'' *''Dicerorhinus yunchuchenensis'' *''Dicerorhinus jeanvireti'' *''Dicerorhinus choukoutienensis'' (synonym of Merck's rhinoceros) *''Dicerorhinus orientalis'' (synonym of Merck's rhinoceros) *''Dicerorhinus nipponicus'' Transferred to ''Dihoplus'' *''Dicerorhinus megarhinus'' *''Dicerorhinus schleiermacheri'' *''Dicero ...
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Dihoplus
''Dihoplus'' is an extinct genus of rhinoceros that lived in Eurasia from the Late Miocene to Early Pleistocene. They were moderately large rhinoceros, with two horns and large, thick nasal bones. Members of ''Dihoplus'' were long placed in ''Dicerorhinus'' (which contains the living Sumatran rhinoceros). Sometimes these species are placed in the related ''Stephanorhinus''. The genus is now generally considered distinct, though there is still debate as to which species should be included; for example, Deng (2011) listed Merck's rhinoceros ''(Stephanorhinus kirchbergensis)'' under ''Dihoplus''. References

Miocene rhinoceroses Pliocene rhinoceroses Pleistocene rhinoceroses Miocene mammals of Europe Miocene mammals of Asia Pliocene mammals of Europe Pliocene mammals of Asia Pleistocene mammals of Europe Pleistocene mammals of Asia {{Paleo-oddtoedungulate-stub ...
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Franz Toula
Franz Toula (20 December 1845 in Vienna – 3 January 1920 in Vienna) was an Austrian geologist, mineralogist and paleontologist. Beginning in 1863 he studied at the Vienna University of Technology, Polytechnic Institute in Vienna (later on, known as ''Technische Hochschule Wien''), where he was a pupil and assistant to Ferdinand von Hochstetter. Later on, he worked for several years as a gymnasium instructor in Vienna. In 1880 he became an associate professor at the ''Technische Hochschule'', where from 1884 to 1917 he served as a full professor of mineralogy and geology.Toula, Franz Edler von (1845–1920), Geologe, Paläontologe und Mineraloge
Österreichisches Biographisches Lexikon und biographische Dokumentation
In 1893/94 he served as rector (academia), ...
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