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Stangeriaceae
Stangeriaceae is the smallest family of the cycads, including only two extant genera, '' Stangeria'' and ''Bowenia'', which both have an underground stem. Taxonomy Although the family was first published by Schimper and Schenk as Stangerieae, the name Stangeriaceae was used by Johnson (who was unaware of previous uses) to include only the genus ''Stangeria.'' Later, Stevenson expanded the family to include ''Bowenia.'' This decision was based on some shared characters (synapomorphies) such as the presence of fused vacularized stipules, the lack or irregular production of cataphylls, and a few other anatomical and morphological traits. Later molecular evidence suggested that the two genera of the Stangeriaceae belonged in different positions within the Zamiaceae, thus leading other authors to propose dissolving the family. Genera * Subfamily Stangerioideae *** '' Stangeria '' Moore (1 species) * Subfamily Bowenioideae *** ''Bowenia'' Hooker ex Hooker f. (2 species) *** † ...
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Stangeria
''Stangeria eriopus'' is a cycad endemic to southern Africa. It is the sole species in the genus ''Stangeria'', most closely related to the Australian genus '' Bowenia'', with which it forms the family Stangeriaceae. Description ''Stangeria eriopus'' is a very long-lived, perennial, evergreen cycad. The stalked, feathered, fern-like leaves are between 25 centimeters and two meters long, with the petiole comprising one third to one half of the overall length (in both varieties). They are pinnately-veined, which distinguishes the species from all other cycads. The petiole comprises half the length of the leaf. The young leaves are bent in bud position, the tip appears rolled up. Young leaves are dotted with short, gray hairs (trichomes), which usually fall off quickly and only stick to the petiole. These trichomes are unbranched and transparent or colored. The species occurs as two variable forms or varieties. The forest form, growing in regions with higher rainfall, is characte ...
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Stangerioideae
''Stangeria eriopus'' is a cycad endemic to southern Africa. It is the sole species in the genus ''Stangeria'', most closely related to the Australian genus ''Bowenia'', with which it forms the family Stangeriaceae. Description ''Stangeria eriopus'' is a very long-lived, perennial, evergreen cycad. The stalked, feathered, fern-like leaves are between 25 centimeters and two meters long, with the petiole comprising one third to one half of the overall length (in both varieties). They are pinnately-veined, which distinguishes the species from all other cycads. The petiole comprises half the length of the leaf. The young leaves are bent in bud position, the tip appears rolled up. Young leaves are dotted with short, gray hairs (trichomes), which usually fall off quickly and only stick to the petiole. These trichomes are unbranched and transparent or colored. The species occurs as two variable forms or varieties. The forest form, growing in regions with higher rainfall, is characterize ...
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Stangeria
''Stangeria eriopus'' is a cycad endemic to southern Africa. It is the sole species in the genus ''Stangeria'', most closely related to the Australian genus ''Bowenia'', with which it forms the family Stangeriaceae. Description ''Stangeria eriopus'' is a very long-lived, perennial, evergreen cycad. The stalked, feathered, fern-like leaves are between 25 centimeters and two meters long, with the petiole comprising one third to one half of the overall length (in both varieties). They are pinnately-veined, which distinguishes the species from all other cycads. The petiole comprises half the length of the leaf. The young leaves are bent in bud position, the tip appears rolled up. Young leaves are dotted with short, gray hairs (trichomes), which usually fall off quickly and only stick to the petiole. These trichomes are unbranched and transparent or colored. The species occurs as two variable forms or varieties. The forest form, growing in regions with higher rainfall, is characterize ...
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Cycads
Cycads are seed plants that typically have a stout and woody (ligneous) trunk with a crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants of a species are either male or female. Cycads vary in size from having trunks only a few centimeters to several meters tall. They typically grow very slowly and live very long. Because of their superficial resemblance, they are sometimes mistaken for palms or ferns, but they are not closely related to either group. Cycads are gymnosperms (naked-seeded), meaning their unfertilized seeds are open to the air to be directly fertilized by pollination, as contrasted with angiosperms, which have enclosed seeds with more complex fertilization arrangements. Cycads have very specialized pollinators, usually a specific species of beetle. Both male and female cycads bear cones (strobili), somewhat similar to conifer cones. Cycads have been reported to fix nitrogen in associatio ...
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Cycad
Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants of a species are either male or female. Cycads vary in size from having trunks only a few centimeters to several meters tall. They typically grow very slowly and live very long. Because of their superficial resemblance, they are sometimes mistaken for Arecaceae, palms or ferns, but they are not closely related to either group. Cycads are gymnosperms (naked-seeded), meaning their fertilization, unfertilized seeds are open to the air to be directly fertilized by pollination, as contrasted with angiosperms, which have enclosed seeds with more complex fertilization arrangements. Cycads have very specialized pollinators, usually a specific species of beetle. Both male and female cycads bear cones (strobilus, strobili), somewhat similar to conife ...
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Bowenia
The genus ''Bowenia'' includes two living and two fossil species of cycads in the family Stangeriaceae, sometimes placed in their own family Boweniaceae. They are entirely restricted to Australia. The two living species occur in Queensland. ''B. spectabilis'' grows in warm, wet, tropical rainforests, on protected slopes and near streams, primarily in the lowlands of the Wet Tropics Bioregion. However, it has a local form with serrate pinna margins that grows in rainforest, ''Acacia''-dominated transition forest, and also ''Casuarina''-dominated sclerophyll forest on the Atherton Tableland, where it is subject to periodic bushfire. ''B. serrulata'' grows in sclerophyll forest and transition forest close to the Tropic of Capricorn. Species The fossil species ''Bowenia eocenica'' is known from deposits in a coal mine in Victoria, Australia, and ''B. papillosa'' is known from deposits in New South Wales. Both fossils are of Eocene age, and consist of leaf A lea ...
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Bowenioideae
The genus ''Bowenia'' includes two living and two fossil species of cycads in the family Stangeriaceae, sometimes placed in their own family Boweniaceae. They are entirely restricted to Australia. The two living species occur in Queensland. ''B. spectabilis'' grows in warm, wet, tropical rainforests, on protected slopes and near streams, primarily in the lowlands of the Wet Tropics Bioregion. However, it has a local form with serrate pinna margins that grows in rainforest, ''Acacia''-dominated transition forest, and also ''Casuarina''-dominated sclerophyll forest on the Atherton Tableland, where it is subject to periodic bushfire. ''B. serrulata'' grows in sclerophyll forest and transition forest close to the Tropic of Capricorn. Species The fossil species ''Bowenia eocenica'' is known from deposits in a coal mine in Victoria, Australia, and ''B. papillosa'' is known from deposits in New South Wales. Both fossils are of Eocene age, and consist of leaf A leaf ( : ...
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Synapomorphy And Apomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis Th ...
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Anfiteatro De Ticó Formation
The Anfiteatro de Ticó Formation is a geological formation from the Deseado Massif in the province of Santa Cruz, Argentina. Paleobiology The Anfiteatro de Ticó preserves a rich and diverse flora including macrofossils, mesofossils, and microfossils. These have been object of detailed investigations since the 1960'. Depositional environments The formation is of continental origin, and includes riparian and lacustrine deposits. Fossil content The following fossils were reported from the formation:Anfiteatro de Ticó Formation
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Flora


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Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, smaller islands. With an area of , Australia is the largest country by area in Oceania and the world's List of countries and dependencies by area, sixth-largest country. Australia is the oldest, flattest, and driest inhabited continent, with the least fertile soils. It is a Megadiverse countries, megadiverse country, and its size gives it a wide variety of landscapes and climates, with Deserts of Australia, deserts in the centre, tropical Forests of Australia, rainforests in the north-east, and List of mountains in Australia, mountain ranges in the south-east. The ancestors of Aboriginal Australians began arriving from south east Asia approximately Early human migrations#Nearby Oceania, 65,000 years ago, during the Last Glacial Period, last i ...
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Queensland
) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_date = Colony of Queensland , established_title2 = Separation from New South Wales , established_date2 = 6 June 1859 , established_title3 = Federation , established_date3 = 1 January 1901 , named_for = Queen Victoria , demonym = , capital = Brisbane , largest_city = capital , coordinates = , admin_center_type = Administration , admin_center = 77 local government areas , leader_title1 = Monarch , leader_name1 = Charles III , leader_title2 = Governor , leader_name2 = Jeannette Young , leader_title3 = Premier , leader_name3 = Annastacia Palaszczuk ( ALP) , legislature = Parliament of Queensland , judiciary = Supreme Court of Queensland , national_representation = Parliament of Australia , national_representation_type ...
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Eastern Cape
The Eastern Cape is one of the provinces of South Africa. Its capital is Bhisho, but its two largest cities are East London and Gqeberha. The second largest province in the country (at 168,966 km2) after Northern Cape, it was formed in 1994 out of the Xhosa homelands or bantustans of Transkei and Ciskei, together with the eastern portion of the Cape Province. The central and eastern part of the province is the traditional home of the indigenous Xhosa people. In 1820 this area which was known as the Xhosa Kingdom began to be settled by Europeans who originally came from England and some from Scotland and Ireland. Since South Africa's early years, many Xhosas believed in Africanism and figures such as Walter Rubusana believed that the rights of Xhosa people and Africans in general, could not be protected unless Africans mobilized and worked together. As a result, the Eastern Cape is home to many anti-apartheid leaders such as Robert Sobukwe, Oliver Tambo, Nelson Mandel ...
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