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SAR Supergroup
SAR is a highly diverse clade of eukaryotes, often considered a supergroup, that includes stramenopiles (heterokonts), alveolates, and rhizarians. It is a node-based taxon (under the Sar name), including all descendants of the three groups' last common ancestor, and comprises most of the now-rejected Chromalveolata. Their sister group has been found to be telonemids, with which they make up the TSAR clade. Harosa is sometimes used synonymously with TSAR. Etymology The name SAR is an acronym derived from the first letters of its three constituent clades; it has been alternatively spelled RAS. The term Harosa (at the subkingdom level) has also been used, with Stramenopiles replaced by its synonym Heterokonta in this variant of the acronym. History of discovery Before the discovery of the SAR supergroup, stramenopiles and alveolates were classified in the supergroup Chromalveolata alongside haptophytes and cryptomonads, being believed to have acquired plastids th ...
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Halvaria
Halvaria is a taxonomic grouping of protists that includes Alveolata and Stramenopiles (Heterokonta). Analyses in 2007 and 2008 revealed that the Stramenopiles and the Alveolata are related, and form a reduced clade of what were seen to be a paraphyletic group, the chromalveolates. The two clades together with the Rhizaria (originally one of the six major eukaryote groups) form a clade dubbed the SAR supergroup. A phylogenomic analysis from 2016 cast doubt on Halvaria, suggesting that Alveolata is the sister group to Rhizaria (making the ''R + A clade'') through new rhizarian sequence data, and that support for Halvaria might be an artifact of low taxon sampling as well as long branch attraction In phylogenetics, long branch attraction (LBA) is a form of systematic error whereby distantly related lineages are incorrectly inferred to be closely related. LBA arises when the amount of molecular or morphological change accumulated within a lin .... However, later analyses from ...
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Synonym (taxonomy)
In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The Botanical nomenclature, botanical and Zoological nomenclature, zoological codes of nomenclature treat the concept of synonymy differently. * In nomenclature, botanical nomenclature, a synonym is a Binomial nomenclature, scientific name that applies to a taxon that now goes by a different scientific name. For example, Carl Linnaeus, Linnaeus was the first to give a scientific name (under the currently used system of scientific nomenclature) to the Norway spruce, which he called ''Pinus abies''. This name is no longer in use, so it is now a synonym of the current scientific name, ''Picea abies''. * In zoology, moving a species from one genus to another results in a different Binomial nomenclature, binomen, but the name is considered an alternative combination rather than a synonym. The concept of synonymy in zoology is reserved f ...
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Chlorarachniophyte
The chlorarachniophytes are a small group of exclusively marine algae widely distributed in tropical and temperate waters. They are typically mixotrophic, ingesting bacteria and smaller protists as well as conducting photosynthesis. Normally they have the form of small amoebae, with branching cytoplasmic extensions that capture prey and connect the cells together, forming a net. These extensions are dependent on the presence of light and polymerization of the actin cytoskeleton. They may also form flagellate zoospores, which characteristically have a single subapical flagellum that spirals backwards around the cell body, and walled coccoid cells. The chloroplasts were presumably acquired by ingesting some green alga. They are surrounded by four membranes, the outermost of which is continuous with the endoplasmic reticulum, and contain a small nucleomorph between the middle two, which is a remnant of the alga's nucleus. This contains a small amount of DNA and divides without for ...
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Endosymbiont
An endosymbiont or endobiont is an organism that lives within the body or cells of another organism. Typically the two organisms are in a mutualism (biology), mutualistic relationship. Examples are nitrogen-fixing bacteria (called rhizobia), which live in the root nodules of legumes, single-cell algae inside Coral reef, reef-building corals, and bacterial endosymbionts that provide essential nutrients to insects. Endosymbiosis played key roles in the development of eukaryotes and plants. Roughly 2.2 billion years ago an archaeon absorbed a bacterium through phagocytosis, that eventually became the mitochondria that provide energy to almost all living Eukaryote, eukaryotic cells. Approximately 1 billion years ago, some of those cells absorbed cyanobacteria that eventually became chloroplasts, organelles that produce energy from sunlight. Approximately 100 million years ago, a lineage of amoeba in the genus ''Paulinella'' independently engulfed a cyanobacterium that evolved to be f ...
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Kelp Forests
Kelp forests are underwater areas with a high density of kelp, which covers a large part of the world's coastlines. Smaller areas of anchored kelp are called kelp beds. They are recognized as one of the most productive and dynamic ecosystems on Earth. Although algal kelp forest combined with coral reefs only cover 0.1% of Earth's total surface, they account for 0.9% of global primary productivity. Kelp forests occur worldwide throughout temperate and polar coastal oceans. In 2007, kelp forests were also discovered in tropical waters near Ecuador. Physically formed by brown macroalgae, kelp forests provide a unique habitat for marine organisms and are a source for understanding many ecological processes. Over the last century, they have been the focus of extensive research, particularly in trophic ecology, and continue to provoke important ideas that are relevant beyond this unique ecosystem. For example, kelp forests can influence coastal oceanographic patterns and provide ma ...
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Phytoplankton
Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater Aquatic ecosystem, ecosystems. The name comes from the Greek language, Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'. Phytoplankton obtain their energy through photosynthesis, as trees and other plants do on land. This means phytoplankton must have light from the sun, so they live in the well-lit surface layers (euphotic zone) of oceans and lakes. In comparison with terrestrial plants, phytoplankton are distributed over a larger surface area, are exposed to less seasonal variation and have markedly faster turnover rates than trees (days versus decades). As a result, phytoplankton respond rapidly on a global scale to climate variations. Phytoplankton form the base of marine and freshwater food webs and are key players in the global carbon cycle. They account for about half of global photosynthetic activity and at least half o ...
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Zooplankton
Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequently, they drift or are carried along by currents in the ocean, or by currents in seas, lakes or rivers. Zooplankton can be contrasted with phytoplankton (cyanobacteria and microalgae), which are the plant-like component of the plankton community (the " phyto-" prefix comes from , although taxonomically ''not'' plants). Zooplankton are heterotrophic (other-feeding), whereas phytoplankton are autotrophic (self-feeding), often generating biological energy and macromolecules through chlorophyllic carbon fixation using sunlightin other words, zooplankton cannot manufacture their own food, while phytoplankton can. As a result, zooplankton must acquire nutrients by feeding on other organisms such as phytoplankton, which are generally smaller t ...
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Hacrobia
The cryptomonads-haptophytes assemblage is a proposed but disputed monophyletic grouping of unicellular eukaryotes that are not included in the SAR supergroup. Several alternative names have been used for the group, including Hacrobia (derived from "ha-" referring to Haptophyta, "-cr-" referring to cryptomonads, and "-bia" as a general suffix referring to life); CCTH (standing for Cryptophyta, Centrohelida, Telonemia and Haptophyta); and "Eukaryomonadae". , it is unclear whether this group is monophyletic or not; results of phylogenetic studies are "often dependent on the selection of taxa and gene data set". Two 2012 studies produced opposite results. Members In the past, heterokonts, haptophytes, and cryptomonads have sometimes been grouped together in a group known as chromists. Though the heterokonts are now split out, Cryptophyta and Haptophyta are considered in some studies to be closely related (and are sometimes simply referred to as the "Cryptophyta+Haptophyta" g ...
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Phylogenetics
In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data and observed heritable traits of DNA sequences, protein amino acid sequences, and morphology. The results are a phylogenetic tree—a diagram depicting the hypothetical relationships among the organisms, reflecting their inferred evolutionary history. The tips of a phylogenetic tree represent the observed entities, which can be living taxa or fossils. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the taxa represented on the tree. An unrooted tree diagram (a network) makes no assumption about directionality of character state transformation, and does not show the origin or "root" of the taxa in question. In addition to their use for inferring phylogenetic pa ...
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Red Algae
Red algae, or Rhodophyta (, ; ), make up one of the oldest groups of eukaryotic algae. The Rhodophyta comprises one of the largest Phylum, phyla of algae, containing over 7,000 recognized species within over 900 Genus, genera amidst ongoing taxonomic revisions. The majority of species (6,793) are Florideophyceae, and mostly consist of multicellular, ocean, marine algae, including many notable seaweeds. Red algae are abundant in marine habitats. Approximately 5% of red algae species occur in freshwater environments, with greater concentrations in warmer areas. Except for two coastal cave dwelling species in the asexual class Cyanidiophyceae, no terrestrial species exist, which may be due to an evolutionary bottleneck in which the last common ancestor lost about 25% of its core genes and much of its evolutionary plasticity. Red algae form a distinct group characterized by eukaryotic cells without flagella and centrioles, chloroplasts without external endoplasmic reticulum or unstack ...
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Symbiogenesis
Symbiogenesis (endosymbiotic theory, or serial endosymbiotic theory) is the leading evolutionary theory of the origin of eukaryotic cells from prokaryotic organisms. The theory holds that mitochondria, plastids such as chloroplasts, and possibly other organelles of eukaryotic cells are descended from formerly free-living prokaryotes (more closely related to the Bacteria than to the Archaea) taken one inside the other in endosymbiosis. Mitochondria appear to be phylogenetically related to Rickettsiales bacteria, while chloroplasts are thought to be related to cyanobacteria. The idea that chloroplasts were originally independent organisms that merged into a symbiotic relationship with other one-celled organisms dates back to the 19th century, when it was espoused by researchers such as Andreas Schimper. The endosymbiotic theory was articulated in 1905 and 1910 by the Russian botanist Konstantin Mereschkowski, and advanced and substantiated with microbiological evidence by Lyn ...
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Cryptomonad
The cryptomonads (or cryptophytes) are a superclass of algae, most of which have plastids. They are traditionally considered a division of algae among phycologists, under the name of Cryptophyta. They are common in freshwater, and also occur in marine and brackish habitats. Each cell is around 10–50 μm in size and flattened in shape, with an anterior groove or pocket. At the edge of the pocket there are typically two slightly unequal flagella. Some may exhibit mixotrophy. They are classified as superclass Cryptomonada, which is divided into two classes: heterotrophic Goniomonadea and phototrophic Cryptophyceae. The two groups are united under three shared morphological characteristics: presence of a periplast, ejectisomes with secondary scroll, and mitochondrial cristae with flat tubules. Genetic studies as early as 1994 also supported the hypothesis that ''Goniomonas'' was sister to Cryptophyceae. A study in 2018 found strong evidence that the common ancestor of Crypto ...
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