Qingtoushan Formation
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Qingtoushan Formation
The Qingtoushan Formation is a Middle Permian-age geologic formation in the Qilian Mountains of Gansu, China. It is known for its diverse tetrapod fauna known as the Dashankou fauna, which likely dates to the Roadian, and includes some of the oldest known therapsids. This formation was previously erroneously named as the Xidagou Formation, a name which applies to otherwise Triassic strata in the northern Qillian Mountains. The formation is over 300 metres thick, and primarily consists of purple-red coarse sandstones, with minor purple mudstone Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from '' shale'' by its lack of fissility (parallel layering).Blatt, H., and R.J. Tracy, 1996, ''Petrology. ....Liu J, Qinghua S, Keqin S, Lu L (2012The horizon of Dashankou Fauna and the Permo-Triassic strata in northern Qilian area, China Vertebr Palasiat 50(4):373–381 (In Chinese) Paleobiota Sy ...
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Geological Formation
A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics ( lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock exposed in a geographical region (the stratigraphic column). It is the fundamental unit of lithostratigraphy, the study of strata or rock layers. A formation must be large enough that it can be mapped at the surface or traced in the subsurface. Formations are otherwise not defined by the thickness of their rock strata, which can vary widely. They are usually, but not universally, tabular in form. They may consist of a single lithology (rock type), or of alternating beds of two or more lithologies, or even a heterogeneous mixture of lithologies, so long as this distinguishes them from adjacent bodies of rock. The concept of a geologic formation goes back to the beginnings of modern scientific geology. The term was used by Abraham Gottlob Wer ...
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Mudstone
Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from '' shale'' by its lack of fissility (parallel layering).Blatt, H., and R.J. Tracy, 1996, ''Petrology.'' New York, New York, W. H. Freeman, 2nd ed, 529 pp. The term ''mudstone'' is also used to describe carbonate rocks (limestone or dolomite) that are composed predominantly of carbonate mud. However, in most contexts, the term refers to siliciclastic mudstone, composed mostly of silicate minerals. The NASA Curiosity rover has found deposits of mudstone on Mars that contain organic substances such as propane, benzene and toluene. Definition There is not a single definition of mudstone that has gained general acceptance,Boggs 2006, p.143 though there is wide agreement that mudstones are fine-grained sedimentary rocks, composed mostly of silicate grains with a grain size less than . Individual grains this size are too small to be disting ...
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Bolosaurid
Bolosauridae is an extinct family of ankyramorph parareptiles known from the latest Carboniferous (Gzhelian) or earliest Permian (Asselian) to the early Guadalupian epoch (latest Roadian stage) of North America, China, Germany, Russia and France. The bolosaurids were unusual for their time period by being bipedal, the oldest known tetrapods to have been so. Their teeth suggest that they were herbivores. The bolosaurids were a rare group and died out without any known descendants. The following cladogram shows the phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... position of the Bolosauridae, from Johannes Müller, Jin-Ling Li and Robert R. Reisz, 2008. References Procolophonomorphs Permian reptiles Paleozoic reptiles of Asia Paleozoic reptiles of Europe Pa ...
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Belebey (genus)
''Belebey'' is an extinct genus of bolosaurid ankyramorph parareptile containing species known from the latest Carboniferous (Gzhelian) or earliest Permian (Asselian) to Guadalupian (Middle Permian) stage of Europe (Russia, France) and Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ... ( China, Qingtoushan Formation). References Procolophonomorphs Permian reptiles of Asia Permian reptiles of Europe Prehistoric reptile genera {{permian-reptile-stub ...
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Raranimus Dashankouensis (2)
''Raranimus'' is an extinct genus of therapsids of the Middle Permian. It was described in 2009 from a partial skull found in 1998 from the Dashankou locality of the Xidagou Formation, outcropping in the Qilian Mountains of Gansu, China. The genus is the most basal known member of the clade Therapsida, to which the later Mammalia belong. Description ''Raranimus'' shares a number of features with later therapsids and ancestral Sphenacodontia. The skull consists of a well preserved rostrum. The teeth suggest a carnivorous lifestyle for ''Raranimus'', as the incisors are recurved and the second canines are serrated on their posterior edges. The incisors are morphologically similar to those seen in more derived theriodonts. The presence of two linguo−labially compressed canines is a diagnostic feature of ''Raranimus''. The presence of two functional canines is characteristic of sphenacodontids, and this condition is seen in no other therapsid other than ''Rananimus''. However, ...
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Raranimus
''Raranimus'' is an extinct genus of therapsids of the Middle Permian. It was described in 2009 from a partial skull found in 1998 from the Dashankou locality of the Xidagou Formation, outcropping in the Qilian Mountains of Gansu, China. The genus is the most basal known member of the clade Therapsida, to which the later Mammalia belong. Description ''Raranimus'' shares a number of features with later therapsids and ancestral Sphenacodontia. The skull consists of a well preserved rostrum. The teeth suggest a carnivorous lifestyle for ''Raranimus'', as the incisors are recurved and the second canines are serrated on their posterior edges. The incisors are morphologically similar to those seen in more derived theriodonts. The presence of two linguo−labially compressed canines is a diagnostic feature of ''Raranimus''. The presence of two functional canines is characteristic of sphenacodontids, and this condition is seen in no other therapsid other than ''Rananimus''. However, the ...
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Stenocybus
''Sinophoneus'' is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 272 to 270 million years ago at the beginning of the Middle Permian (Lower Roadian) in what is now the Gansu Province in northern China. It is known by a skull of an adult individual (the holotype GMV1601), as well as by many skulls of juvenile specimens. The latter were first considered as belonging to a different animal, named ''Stenocybus'', before being reinterpreted as immature ''Sinophoneus''. ''Sinophoneus'' shows a combination of characters present in other anteosaurs. Its bulbous profile snout and external nostrils located in front of the canine are reminiscent of the basal anteosaur ''Archaeosyodon'', while its massive transerse pterygoids processes with enlarged distal ends are more similar to the more derived anteosaurs ''Anteosaurus'' and ''Titanophoneus''. First phylogenetic analyzes identified ''Sinophoneus'' as the most basal Anteosaurinae. A mo ...
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Sinophoneus09
''Sinophoneus'' is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 272 to 270 million years ago at the beginning of the Middle Permian (Lower Roadian) in what is now the Gansu Province in northern China. It is known by a skull of an adult individual (the holotype GMV1601), as well as by many skulls of juvenile specimens. The latter were first considered as belonging to a different animal, named ''Stenocybus'', before being reinterpreted as immature ''Sinophoneus''. ''Sinophoneus'' shows a combination of characters present in other anteosaurs. Its bulbous profile snout and external nostrils located in front of the canine are reminiscent of the basal anteosaur ''Archaeosyodon'', while its massive transerse pterygoids processes with enlarged distal ends are more similar to the more derived anteosaurs ''Anteosaurus'' and ''Titanophoneus''. First phylogenetic analyzes identified ''Sinophoneus'' as the most basal Anteosaurinae. A mo ...
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Dinocephalian
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the Biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They include carniv ...
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Anteosauridae
Anteosauridae is an extinct family of large carnivorous dinocephalian therapsids that are known from the Middle Permian of Asia, Africa, and South America.These animals were by far the largest predators of the Permian period, with skulls reaching 80 cm in length in adult individuals, far larger than the biggest gorgonopsian. Description Anteosaurids are characterized by very large pointed incisors and canines, bulbous spatulate (spoon shaped) postcanines, a very strongly upturned margin of the premaxilla, so the front of mouth curves strongly upwards, and a long, very robust lower jaw (Boonstra 1963). Anteosaurids distinguished from ''Brithopus'' and other non-anteosaurid anteosaurians by the presence of a large thickened region or "boss" on the side of the angle of the lower jaw; this was probably used in intraspecific behaviour. In '' Doliosauriscus'' and ''Anteosaurus'', not only was this boss very prominent, but the bones were very thick and rugose. The same situation ...
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Sinophoneus
''Sinophoneus'' is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 272 to 270 million years ago at the beginning of the Middle Permian (Lower Roadian) in what is now the Gansu Province in northern China. It is known by a skull of an adult individual (the holotype GMV1601), as well as by many skulls of juvenile specimens. The latter were first considered as belonging to a different animal, named ''Stenocybus'', before being reinterpreted as immature ''Sinophoneus''. ''Sinophoneus'' shows a combination of characters present in other anteosaurs. Its bulbous profile snout and external nostrils located in front of the canine are reminiscent of the basal anteosaur ''Archaeosyodon'', while its massive transerse pterygoids processes with enlarged distal ends are more similar to the more derived anteosaurs ''Anteosaurus'' and ''Titanophoneus''. First phylogenetic analyzes identified ''Sinophoneus'' as the most basal Anteosaurinae. A mo ...
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