Pterosaur Phylogeny
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Pterosaur Phylogeny
This phylogeny of pterosaurs entails the various phylogenetic trees used to classify pterosaurs throughout the years and varying views of these animals. Pterosaur phylogeny is currently highly contested and several hypotheses are presented below. Unwin (2003) The matrix includes 19 pterosaur groups (most of which are supra-specific) plus a single outgroup (''Euparkeria capensis''). The taxa were coded for 60 characters. Kellner (2003) The matrix includes 39 valid pterosaur species, although ''Rhamphorhynchus longicaudus'' and ''Nyctosaurus bonneri'' are usually considered to be synonymous with ''R. muensteri'' and ''N. gracilis'' respectively, plus a three outgroup species (''Ornithosuchus longidens'', '' Herrerasaurus ischigualastensis'' and ''Scleromochlus taylori''). The taxa were coded for 74 characters. Andres and Myers (2013) In 2010, Brian Blake Andres wrote a review of pterosaur phylogeny in his dissertation. His phylogenetic analysis combined data mainly from three ...
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Ornithocheiroidea
Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with fossil remains found all over the world except Antarctica. Ornithocheiroids were the most advanced group of pterosaurs, as the group includes the clade Azhdarchoidea, of which its members lived until the Maastrichtian stage of the Late Cretaceous, around 66 million years ago. Notable pterosaurs from this group include the pteranodontians ''Pteranodon'' and ''Nyctosaurus'', the ornithocheirid ''Ornithocheirus'', the anhanguerid ''Tropeognathus'', as well as the azhdarchids ''Hatzegopteryx'' and ''Quetzalcoatlus''. Classification The name Ornithocheiroidea was originally defined as an apomorphy-based taxon by Christopher Bennett in 1994. It was given a relationship-based definition in 2003 by Alexander Kellner, who defined it as the least in ...
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Ctenochasmatidae
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods. The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Classification Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "cha ...
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Lonchodectes
''Lonchodectes'' (meaning " lance biter") was a genus of lonchodectid pterosaur from several formations dating to the Turonian ( Late Cretaceous) of England, mostly in the area around Kent. The species belonging to it had been assigned to '' Ornithocheirus'' until David Unwin's work of the 1990s and 2000s.Kellner, A.W.A. (2003). Pterosaur phylogeny and comments on the evolutionary history of the group: In: Buffetaut, E., and Mazin, J.-M. (Eds.). ''Evolution and Palaeobiology of Pterosaurs''. Geological Society Special Publication 217:105-137. 1-86239-143-2. Several potential species are known; most are based on scrappy remains, and have gone through several other generic assignments. The genus is part of the complex taxonomy issues surrounding Early Cretaceous pterosaurs from Brazil and England, such as '' Amblydectes'', '' Anhanguera'', '' Coloborhynchus'', and ''Ornithocheirus''. History and species Numerous species have been referred to this genus over time, and on ...
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Pterodactylus
''Pterodactylus'' (from Greek () meaning 'winged finger') is an extinct genus of pterosaurs. It is thought to contain only a single species, ''Pterodactylus antiquus'', which was the first pterosaur to be named and identified as a flying reptile. Fossil remains of ''Pterodactylus'' have primarily been found in the Solnhofen limestone of Bavaria, Germany, which dates from the Late Jurassic period (early Tithonian stage), about 150.8 to 148.5 million years ago. More fragmentary remains of ''Pterodactylus'' have tentatively been identified from elsewhere in Europe and in Africa. ''Pterodactylus'' was a generalist carnivore that probably fed on a variety of invertebrates and vertebrates. Like all pterosaurs, ''Pterodactylus'' had wings formed by a skin and muscle membrane stretching from its elongated fourth finger to its hind limbs. It was supported internally by collagen fibres and externally by keratinous ridges. ''Pterodactylus'' was a small pterosaur compared to other famo ...
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Euctenochasmatia
Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of ''Pterodactylus'' and ''Ctenochasma'', and all their descendants. Euctenochasmatians were specialized pterosaurs that had elongated necks as well as specialized teeth. A peculiar family within this group is the Ctenochasmatidae, which most of the members had very distinguishing teeth that were lined within their elongated snouts. A genus called ''Pterofiltrus'' only had 112 teeth, but these teeth cover about 55.8% of the total skull, and the skull itself measured about in length. Description Euctenochasmatians had very distinctive features in comparison to other pterosaurs, including the shape of their jaws, as well as their highly specialized teeth. These teeth are thought to have been used for filter-feeding, the genus ''Pterodaustro'' for example, had a long snout and its lower jaws curve strongly upwards, and th ...
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Gallodactylidae
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey. History Gallodactylidae was named to contain ''Gallodactylus'' (now usually considered a synonym of ''Cycnorhamphus'') and its closest relatives. Many subsequent studies, however, showed that ''Gallodactylus'' did not form a clade with any non-synonymous pterosaurs that were not themselves part of a different family, and so the name was often ignored. The name returned to common use with the discovery of ''Gladocephaloideus'', a Chinese pterosaur species that shared many similarities with ''Cycnorhamphus''. Among other features, the Gallodactylidae was distinguished by ...
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Ctenochasmatoidea
Ctenochasmatoidea is a group of early pterosaurs within the suborder Pterodactyloidea. Their remains are usually found in what were once coastal or lake environments. They generally had long wings, long necks, and highly specialized teeth. Evolutionary history The earliest known ctenochasmatoid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Ecology Most ctenochasmatoids were aquatic or semi-aquatic pterosaurs, possessing large webbed hindfeet and long ...
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Lophocratia
Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoids by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like '' Nyctosaurus'' and '' Tupandactylus''. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles. ...
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Pteranodontinae
''Pteranodon'' (); from Ancient Greek (''pteron'', "wing") and (''anodon'', "toothless") is a genus of pterosaur that included some of the largest known flying reptiles, with ''P. longiceps'' having a wingspan of . They lived during the late Cretaceous geological period of North America in present-day Kansas, Nebraska, Wyoming, South Dakota and Alabama. More fossil specimens of ''Pteranodon'' have been found than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved with nearly complete skulls and articulated skeletons. It was an important part of the animal community in the Western Interior Seaway. ''Pteranodon'' was not a dinosaur. By definition, all dinosaurs belong to the group Dinosauria; ''Pteranodon'' belongs to the group Pterosauria. Nonetheless, ''Pteranodon'' is the most famous pterosaur, frequently featured in dinosaur media and strongly associated with dinosaurs by the general public. While not dinosaurs, pterosau ...
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Nyctosaurus
''Nyctosaurus'' (meaning "night lizard" or "bat lizard") is a genus of nyctosaurid pterosaur from the Late Cretaceous period of what is now the Niobrara Formation of the mid-western United States, which, during the time ''Nyctosaurus'' was alive, was covered in an extensive shallow sea. Some remains belonging to a possible ''Nyctosaurus'' species called ''N.lamegoi'' have been found in Brazil, making ''Nyctosaurus'' more diverse. The genus ''Nyctosaurus'' has had numerous species referred to it, though how many of these may actually be valid requires further study. At least one species possessed an extraordinarily large antler-like cranial crest. ''Nyctosaurus'' was a mid-sized pterosaur that lived along the shores of the Niobrara Formation of the United States, which back then was within a large inland sea called the Western Interior Seaway. It has been suggested that it would have flown similar to modern-day soaring birds such as albatrosses, which consisted of flying very lon ...
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Pteranodontidae
The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head (most famously seen in ''Pteranodon'' itself). The spectacularly-crested ''Nyctosaurus'' is sometimes included in this family, though usually placed in its own family, the Nyctosauridae (Nicholson & Lydekker, 1889). Modern researchers differ in their use of the concept. S. Christopher Bennett and Alexander Kellner have concluded that ''Nyctosaurus'' was not a pteranodontid. In 1994 Bennett defined a clade Pteranodontidae, also including species of the Anhangueridae. However, this definition has not been accepted by other workers. Alexander Kellner, for example, named several additional species for specimens previously classified as ''Pteranodon'', and placed ''P. sternbergi'' in a distinct genus, ''Geosternbergia''. Kellner re-defined Ptera ...
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