Protheriodon
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Protheriodon
''Protheriodon'' is an extinct genus of probainognathian cynodonts which existed in the Santa Maria Formation of the Paraná Basin in southeastern Brazil during the middle Triassic period. It contains the species ''Protheriodon estudianti''. It was first described by Argentine palaeontologist José Bonaparte José Fernando Bonaparte (14 June 1928 – 18 February 2020) was an Argentine paleontologist who discovered a plethora of South American dinosaurs and mentored a new generation of Argentine paleontologists . One of the best-known Argentine paleo ..., who assigned it to the family Brasilodontidae. More recent studies have however recovered it in a more basal position than other brasilodontids, just outside Prozostrodontia. Cladogram from Martinelli ''et al.'', 2017: References Prehistoric cynodont genera Ladinian genera Middle Triassic synapsids of South America Triassic Brazil Fossils of Brazil Santa Maria Formation Fossil taxa described in 2006 Taxa ...
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Brasilodontidae
''Brasilodon'' ("tooth from Brazil") is an extinct genus of small, mammal-like cynodonts that lived in what is now Brazil during the Norian age of the Late Triassic epoch, about 225.42 million years ago. While no complete skeletons have been found, the length of ''Brasilodon'' has been estimated at around . Its dentition shows that it was most likely an insectivore. The genus is monotypic, containing only the species ''B. quadrangularis''. ''Brasilodon'' belongs to the family Brasilodontidae, whose members were some of the closest relatives of mammals, the only cynodonts alive today. Two other brasilodontid genera, ''Brasilitherium'' and ''Minicynodon'', are now considered to be junior synonyms of ''Brasilodon''. Discovery and naming The first three specimens referred to ''Brasilodon quadrangularis'' were found at the Linha São Luiz site, a quarry near the town of Faxinal do Soturno in the state of Rio Grande do Sul. The rocks where ''Brasilodon'' was found belong to the upp ...
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Bonacynodon
''Bonacynodon'' is an extinct genus of cynodonts that lived in what is now southern Brazil during the Triassic period (Ladinian–Carnian ages). The genus is monotypic, containing only the type species ''Bonacynodon schultzi''. ''B. schultzi'' is known from two specimens, consisting of two partial skulls and some badly preserved parts of the postcranium. Both specimens were recovered from the Pinheiros-Chiniquá Sequence, part of the Santa Maria Supersequence of the Paraná Basin. This sequence preserves a faunal association known as the ''Dinodontosaurus'' Assemblage Zone, which contains numerous other species of cynodonts, dicynodonts and reptiles. ''Bonacynodon'' was a small, likely insectivorous cynodont, whose length has been estimated at around . It can be distinguished from other cynodonts by its large, serrated (saw-like) canine teeth. Together with the genus ''Probainognathus'' of Argentina, it made up the family Probainognathidae, one of the earliest-diverging line ...
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Candelariodon
''Candelariodon'' is an extinct genus of carnivorous probainognathian cynodonts from the Middle to Late Triassic Santa Maria Formation of the Paraná Basin in Rio Grande do Sul state, Brazil. ''Candelariodon'' is known from a partial mandible having some complete teeth. It was first named by Téo Veiga de Oliveira, Cesar Leandro Schultz, Marina Bento Soares and Carlos Nunes Rodrigues in 2011 and the type species is ''Candelariodon barberenai''. Description The holotype (MMACR PV-0001-T) of ''Candelariodon barberenai'' comes from the ''Dinodontosaurus'' Assemblage Zone of the Pinheros-Chiniquá Sequence, Santa Maria Supersequence. It corresponds to the lower portion of the traditional Santa Maria Formation and the Santa Maria 1 Sequence of Zerfass et al. (2003). The outcrop that yielded MMACR PV-0001-T is located about south of Candelária, Rio Grande do Sul, southeastern Brazil, in the Pinheiro, an area in which several tetrapods characteristic of the ''Dinodontosauru ...
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Santa Maria Formation
The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age (Late Triassic), and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and other dinosauromorphs, including the herrerasaurid '' Staurikosaurus'', the basal sauropodomorphs ''Buriolestes'' and '' Saturnalia,'' and the lagerpetid ''Ixalerpeton''. The formation is named after the city of Santa Maria in the central region of Rio Grande do Sul, where outcrops were first studied. The Santa Maria Formation makes up the majority of the Santa Maria Supersequence, which extends through the entire Late Triassic. The Santa Maria Supersequence is divided into four geological sequences, separated from each other by short unconformities. The first two of these sequences (Pinheiros-Chiniquá and Santa Cruz sequences) lie entirely within the Santa Maria Formation, while the third (the Candelária sequence) is shared with the ov ...
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Probainognathia
Probainognathia is one of the two major subgroups of the clade Eucynodontia, the other being Cynognathia. The earliest forms were carnivorous and insectivorous, though some groups eventually also evolved herbivorous diets. The earliest and most basal probainognathian is the Middle Triassic (Anisian) aged ''Lumkuia'', from South Africa, though probainognathians would not become prominent until the mid Norian stage of the Late Triassic. Three groups survived the extinction at the end of Triassic: Tritheledontidae and Tritylodontidae, which both survived until the Jurassic—the latter even into the Cretaceous (''Montirictus'' and '' Xenocretosuchus'')—and Mammaliaformes, which includes the mammals. Phylogeny Below is a cladogram from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships: See also * Evolution of mammals * List of prehistoric mammals This is an incomplete list of prehistoric mammals. It does not include extant mam ...
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Probainognathia
Probainognathia is one of the two major subgroups of the clade Eucynodontia, the other being Cynognathia. The earliest forms were carnivorous and insectivorous, though some groups eventually also evolved herbivorous diets. The earliest and most basal probainognathian is the Middle Triassic (Anisian) aged ''Lumkuia'', from South Africa, though probainognathians would not become prominent until the mid Norian stage of the Late Triassic. Three groups survived the extinction at the end of Triassic: Tritheledontidae and Tritylodontidae, which both survived until the Jurassic—the latter even into the Cretaceous (''Montirictus'' and '' Xenocretosuchus'')—and Mammaliaformes, which includes the mammals. Phylogeny Below is a cladogram from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships: See also * Evolution of mammals * List of prehistoric mammals This is an incomplete list of prehistoric mammals. It does not include extant mam ...
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Ladinian
The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic). The Ladinian is coeval with the Falangian regional stage used in China. Stratigraphic definitions The Ladinian was established by Austrian geologist Alexander Bittner in 1892. Its name comes from the Ladin people that live in the Italian Alps (in the Dolomites, then part of Austria-Hungary). The base of the Ladinian Stage is defined as the place in the stratigraphic record where the ammonite species '' Eoprotrachyceras curionii'' first appears or the first appearance of the conodont ''Budurovignathus praehungaricus''. The global reference profile for the base (the GSSP) is at an outcrop in the river bed of the Caffaro river at Bagolino, in the province of Brescia, northern Italy.The GSSP was established by Brack ''et al.'' (2005) The t ...
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Fossil Taxa Described In 2006
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolute ...
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Fossils Of Brazil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolut ...
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Triassic Brazil
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic. The Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Three categories of organisms can be distinguished in the Triassic record: survivors from the extinction event, new groups that flourished briefly, and other new groups that went on to dominate the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized subgroup of archosaur ...
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Middle Triassic Synapsids Of South America
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Ladinian Genera
The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic). The Ladinian is coeval with the Falangian regional stage used in China. Stratigraphic definitions The Ladinian was established by Austrian geologist Alexander Bittner in 1892. Its name comes from the Ladin people that live in the Italian Alps (in the Dolomites, then part of Austria-Hungary). The base of the Ladinian Stage is defined as the place in the stratigraphic record where the ammonite species '' Eoprotrachyceras curionii'' first appears or the first appearance of the conodont ''Budurovignathus praehungaricus''. The global reference profile for the base (the GSSP) is at an outcrop in the river bed of the Caffaro river at Bagolino, in the province of Brescia, northern Italy.The GSSP was established by Brack ''et al.'' (2005) The ...
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