Proterochampsians
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Proterochampsians
Proterochampsia is a clade of early archosauriform reptiles from the Triassic period. It includes the Proterochampsidae (e.g. '' Proterochampsa'', '' Chanaresuchus'' and '' Tropidosuchus'') and probably also the Doswelliidae. Nesbitt (2011) defines Proterochampsia as a stem-based taxon that includes '' Proterochampsa'' and all forms more closely related to it than ''Euparkeria'', ''Erythrosuchus'', ''Passer domesticus'' (the House Sparrow), or ''Crocodylus niloticus'' (the Nile crocodile). Therefore, the inclusion of Doswelliidae in it is dependent upon whether ''Doswellia'' and '' Proterochampsa'' form a monophyletic group to the exclusion of Archosauria and other related groups. Description Nesbitt (2011) found that Proterochampsians share several distinguishing characteristics, or synapomorphies. A prominent ridge runs along the length of the jugal, a bone below the eye. Another ridge is present on the quadratojugal, a bone positioned toward the back of the skull behind the j ...
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Tropidosuchus
''Tropidosuchus'' is an extinct genus of carnivorous archosauriforms from the Middle Triassic period (Anisian to Ladinian stage). It is a proterochampsid which lived in what is now Argentina. It is known from the holotype PVL 4601, which consists of partial skeleton. It was found in the Chañares Formation and its type locality is the Chañares River. It was first named by A. B. Arcucci in 1990 and the type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ... is ''Tropidosuchus romeri''. References Proterochampsians Ladinian genera Middle Triassic reptiles of South America Triassic Argentina Fossils of Argentina Chañares Formation Fossil taxa described in 1990 Prehistoric reptile genera {{triassic-reptile-stub ...
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Pseudochampsa Ischigualastensis
''Pseudochampsa'' is an extinct genus of proterochampsid archosauriform known from the Late Triassic (Carnian) Cancha de Bochas Member of the Ischigualasto Formation of San Juan Province, Ischigualasto-Villa Unión Basin in northwestern Argentina. It contains a single species, ''Pseudochampsa ischigualastensis'', originally named as a second species of the closely related '' Chanaresuchus'', based on a fairly complete articulated skeleton and skull.Trotteyn et al., 2012 A revision of the remains concluded that it was best to move to species to its own genus, as no traits were found to unite ''P. ischigualastensis'' and the type species of ''Chanaresuchus'' to the exclusion of other proterochampsids. A phylogenetic analysis places both species in a polytomy with '' Gualosuchus'' as the most advanced members of Proterochampsia.Trotteyn & Ezcurra, 2014 Discovery ''Pseudochampsa'' is known solely from the holotype PVSJ 567, a nearly complete and articulated individual hous ...
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Doswelliidae
Doswelliidae is an extinct family of carnivorous archosauriform reptiles that lived in North America and Europe during the Middle to Late Triassic period. Long represented solely by the heavily-armored reptile ''Doswellia'', the family's composition has expanded since 2011, although two supposed South American doswelliids ('' Archeopelta'' and '' Tarjadia'') were later redescribed as erpetosuchids. Doswelliids were not true archosaurs, but they were close relatives and some studies have considered them among the most derived non-archosaurian archosauriforms. They may have also been related to the Proterochampsidae, a South American family of crocodile-like archosauriforms. Description Doswelliids are believed to be semiaquatic carnivores similar to crocodilians in appearance, as evidenced by their short legs and eyes and nostrils which are set high on the head, though the putative member '' Scleromochlus'' has been interpreted as a frog-like hopper by one study. They had ...
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Proterochampsidae
Proterochampsidae is a family of proterochampsian archosauriforms. Proterochampsids may have filled an ecological niche similar to modern crocodiles, and had a general crocodile-like appearance. They lived in what is now South America in the Middle and Late Triassic. Description Proterochampsids have long, crocodile-like skulls. The posterior portion of the skull is wide while the snout is very narrow. Most proterochampsids also have downturned snouts. Like many early archosauriforms, they also have dermal armour. Proterochampsids have small holes called dorsal fenestrae at the top of their skulls. Unlike other early archosauromorphs, they do not have a parietal foramin, which in many reptiles holds a parietal eye. The postorbital bones behind the eye sockets have thick, jagged crests. As another diagnostic feature of the group, the holes that allow the passage of the internal carotid artery through the braincase open at the sides of a bony projection called the basipterygoid p ...
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Proterochampsa BW
''Proterochampsa'' is a genus of proterochampsid archosauromorph from the Late Triassic (Carnian) Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina and the Alemoa Member of the Santa Maria Formation in the Paraná Basin in southeastern Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area .... Average length of skull of ''P. barrionuevoi'' is approximately . References Further reading * F. Abdala, M. C. Barberena, and J. Dornelles. 2002. A new species of the traversodontid cynodont Exaeretodon from the Santa Maria Formation (Middle/Late Triassic) of southern Brazil. Journal of Vertebrate Paleontology 22(2):313-325 * * O. A. Reig. 1963. La presencia de dinosaurios saurisquios en los "Estratos de Ischigu ...
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Doswellia
''Doswellia'' is an extinct genus of archosauriform from the Late Triassic of North America. It is the most notable member of the family Doswelliidae, related to the proterochampsids. ''Doswellia'' was a low and heavily built carnivore which lived during the Carnian stage of the Late Triassic. It possesses many unusual features including a wide, flattened head with narrow jaws and a box-like rib cage surrounded by many rows of bony plates. The type species ''Doswellia kaltenbachi'' was named in 1980 from fossils found within the Vinita member of the Doswell Formation (formerly known as the Falling Creek Formation) in Virginia. The formation, which is found in the Taylorsville Basin, is part of the larger Newark Supergroup. ''Doswellia'' is named after Doswell, the town from which much of the taxon's remains have been found. A second species, ''D. sixmilensis,'' was described in 2012 from the Bluewater Creek Formation of the Chinle Group in New Mexico; however, this species was sub ...
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Proterochampsa
''Proterochampsa'' is a genus of proterochampsid archosauromorph from the Late Triassic (Carnian) Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina and the Alemoa Member of the Santa Maria Formation in the Paraná Basin in southeastern Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area .... Average length of skull of ''P. barrionuevoi'' is approximately . References Further reading * F. Abdala, M. C. Barberena, and J. Dornelles. 2002. A new species of the traversodontid cynodont Exaeretodon from the Santa Maria Formation (Middle/Late Triassic) of southern Brazil. Journal of Vertebrate Paleontology 22(2):313-325 * * O. A. Reig. 1963. La presencia de dinosaurios saurisquios en los "Estratos de Ischig ...
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Chanaresuchus
''Chanaresuchus'' is an extinct genus of proterochampsian archosauriform. It was of modest size for a proterochampsian, being on average just over a meter in length. Fossils are known from the Middle and Late Triassic of La Rioja Province, Argentina and Rio Grande do Sul (geoparque Paleorrota), Brazil. The type species and only currently known species is ''Chanaresuchus bonapartei'' was named from the Ladinian-age Chañares Formation in 1971. A second species ''C. ischigualastensis'' named in 2012 from the late Carnian-age Ischigualasto Formation, was briefly assigned to ''Chanaresuchus'' before being moved to its own genus '' Pseudochampsa'' in 2014. ''C. bonapartei'' has recently been found in the Carnian Santa Maria Formation in Brazil. ''Chanaresuchus'' appears to be one of the most common archosauriforms from the Chanares Formation due to the abundance of specimens referred to the genus. Much of the material has been found by the La Plata-Harvard expedition of 1964-65. ''Cha ...
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Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and Ma (million years ago). It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages. Formerly the middle series in the Triassic was also known as Muschelkalk. This name is now only used for a specific unit of rock strata with approximately Middle Triassic age, found in western Europe. Middle Triassic fauna Following the Permian–Triassic extinction event, the most devastating of all mass-extinctions, life recovered slowly. In the Middle Triassic, many groups of organisms reached higher diversity again, such as the marine reptiles (e.g. ichthyosaurs, sauropterygians, thallatosaurs), ray-finned fish and many invertebrate groups like ...
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Synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Jugal
The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy The jugal bone is located on either side of the skull in the circumorbital region. It is the origin of several masticatory muscles in the skull. The jugal and lacrimal bones are the only two remaining from the ancestral circumorbital series: the prefrontal, postfrontal, postorbital, jugal, and lacrimal bones. During development, the jugal bone originates from dermal bone. In dinosaurs This bone is considered key in the determination of general traits in cases in which the entire skull has not been found intact (for instance, as with dinosaurs in paleontology). In some dinosaur genera the jugal also forms part of the lower margin of either the antorbital fenestra or the infratemporal fenestra, or both. Most commonly, this bone articu ...
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Quadratojugal
The quadratojugal is a skull bone present in many vertebrates, including some living reptiles and amphibians. Anatomy and function In animals with a quadratojugal bone, it is typically found connected to the jugal (cheek) bone from the front and the squamosal bone from above. It is usually positioned at the rear lower corner of the cranium. Many modern tetrapods lack a quadratojugal bone as it has been lost or fused to other bones. Modern examples of tetrapods without a quadratojugal include salamanders, mammals, birds, and squamates (lizards and snakes). In tetrapods with a quadratojugal bone, it often forms a portion of the jaw joint. Developmentally, the quadratojugal bone is a dermal bone in the temporal series, forming the original braincase. The squamosal and quadratojugal bones together form the cheek region and may provide muscular attachments for facial muscles. In reptiles and amphibians In most modern reptiles and amphibians, the quadratojugal is a prominent, strapl ...
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