Protaspididae
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Protaspididae
Protaspididae is an extinct family of pteraspidid heterostracan agnathans. Fossils of the various genera are found in early Devonian-aged marine strata. Protaspidids were once thought to represent a transitional form between the Pteraspididae and the Psammosteida, bearing the broad head shield shape of the latter, due to a more benthic (bottom-dwelling) existence, but recent phylogenical comparisons demonstrate that the protaspidids are actually highly derived pteraspidids,Pernegre, Vincent N., and David K. Elliott. "Phylogeny of the Pteraspidiformes (Heterostraci), Silurian–Devonian jawless vertebrates." Zoologica Scripta 37.4 (2008): 391-403. and that the anchipteraspidids, the most primitive of pteraspidids, are the sister-group of the Psammosteids. Genera Many genera are found in Lower Devonian marine deposits of the United States, especially of Utah and Wyoming. Others are found in Lower Devonian marine strata of Europe, especially of Svalbard, Norway, and Podolia, Ukrai ...
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Pteraspidida
Pteraspidiformes is an extinct order of heterostracan agnathan vertebrates known from extensive fossil remains primarily from Early Devonian strata of Europe and North America, and from Upper Silurian Canada. Anatomy A pteraspidiform heterostracan has the cephalothorax enclosed in armor, formed from several plates, including dorsal, ventral, rostral, pineal plates, a dorsal spine derived from a scale, and a large, scale-covered tail. Many genera were benthic, others were apparently active swimming nekton.Botella, Hector, and Richard A. Farina. "Flow pattern around the rigid cephalic shield of the Devonian agnathan Errivaspis waynensis (Pteraspidiformes: Heterostraci)." Palaeontology 51.5 (2008): 1141-1150/ref> Delicate, finger-like components of the anterior end of the ventral plate forming the edges of the mouth suggest that pteraspidiform heterostracans were filter-feeding, filter-feeders that selectively filtered specific sized plankton from the water column.Purnell, Mark ...
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Heterostraci
Heterostraci (Ancient Greek, ἕτερος+ὄστρακον "those itha different shell" i is pl. of -us is an extinct subclass of pteraspidomorph jawless vertebrate that lived primarily in marine and estuary environments. Heterostraci existed from the mid-Ordovician to the conclusion of the Devonian. Description and anatomy The Heterostracans differed from other Paleozoic agnathan taxa both in the arrangement and histology of their scales. Most heterostracans had two plates which form a large dorsal shield and a large ventral shield, and had series of scales arranged in various patterns on the sides of their bodies, the exact pattern differing from one group to another. In a few primitive forms, such as '' Lepidaspis'', the dorsal and ventral shields are composed of a mosaic of tiny scales. In most other known forms, though, these tiny scales have fused together to form the shield-plates. The scales of heterostracans are histologically distinct from other vertebrate ...
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Protopteraspididae
Protopteraspididae is an extinct family of pteraspidid heterostracan agnathans. Fossils of the various genera are found in early Devonian-aged marine strata. Protopteraspidids were once thought to represent a taxon of basal pteraspidids (hence "proto") but recent evaluations demonstrate that Protopteraspididae is a paraphyletic group of various transitional forms representing a gradual transition between the more advanced Pteraspoidei (comprising Pteraspididae, '' Gigantaspis'' and Protaspididae),Pernegre, Vincent N., and David K. Elliott. "Phylogeny of the Pteraspidiformes (Heterostraci), Silurian–Devonian jawless vertebrates." Zoologica Scripta 37.4 (2008): 391-403. and the anchipteraspidids and the Psammosteids. Genera Protopteraspids are found in Lower Devonian marine strata of the Western United States, Northern Canada, England, France, Belgium, Svalbard, Norway, and Podolia, Ukraine.Blieck, Alain. "Les hétérostracés Ptéraspidiformes, agnathes du Silurien-Dévonien ...
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Nevada
Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the List of U.S. states and territories by area, 7th-most extensive, the List of U.S. states and territories by population, 32nd-most populous, and the List of U.S. states and territories by population density, 9th-least densely populated of the U.S. states. Nearly three-quarters of Nevada's people live in Clark County, Nevada, Clark County, which contains the Las Vegas–Paradise, NV MSA, Las Vegas–Paradise metropolitan area, including three of the state's four largest incorporated cities. Nevada's capital is Carson City, Nevada, Carson City. Las Vegas is the largest city in the state. Nevada is officially known as the "Silver State" because of the importance of silver to its history and economy. It is also known as the "Battle ...
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Psammosteida
Psammosteida also called as Psammosteoidei is a suborder of pteraspidid heterostracan agnathans. The psammosteids had broad, flattened bodies, suggesting a predominantly benthic habit. The earliest unequivocal psammosteid is '' Drepanaspis'' of Early Devonian Germany, which is either included in the family Psammosteidae, or placed within its own family, Drepanaspididae. If the late Silurian/Early Devonian '' Weigeltaspis'' is a psammosteid, as opposed to being a traquairaspid, then that genus, instead, would be the oldest psammosteid. However, its placement within Heterostraci remains a matter of debate. Other notable psammosteids include '' Psammosteus'', and '' Obruchevia'', two genera of enormous species with dorsal shields around one meter in diameter. The Psammosteids were the only heterostracans to survive to the end of the Devonian, where they finally perish during the Hangenberg event The Hangenberg event, also known as the Hangenberg crisis or end-Devonian extinct ...
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Ukraine
Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian invasion, it was the eighth-most populous country in Europe, with a population of around 41 million people. It is also bordered by Belarus to the north; by Poland, Slovakia, and Hungary to the west; and by Romania and Moldova to the southwest; with a coastline along the Black Sea and the Sea of Azov to the south and southeast. Kyiv is the nation's capital and largest city. Ukraine's state language is Ukrainian; Russian is also widely spoken, especially in the east and south. During the Middle Ages, Ukraine was the site of early Slavic expansion and the area later became a key centre of East Slavic culture under the state of Kievan Rus', which emerged in the 9th century. The state eventually disintegrated into rival regional po ...
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Podolia
Podolia or Podilia ( uk, Поділля, Podillia, ; russian: Подолье, Podolye; ro, Podolia; pl, Podole; german: Podolien; be, Падолле, Padollie; lt, Podolė), is a historic region in Eastern Europe, located in the west-central and south-western parts of Ukraine and in northeastern Moldova (i.e. northern Transnistria). The name derives from Old Slavic ''po'', meaning "by/next to/along" and ''dol'', "valley" (see dale). Geography The area is part of the vast East European Plain, confined by the Dniester River and the Carpathian arc in the southwest. It comprises an area of about , extending for from northwest to southeast on the left bank of the Dniester. In the same direction run two ranges of relatively low hills separated by the Southern Bug, ramifications of the Avratynsk heights. The Podolian Upland, an elongated, up to high plateau stretches from the Western and Southern Bug rivers to the Dniester, and includes hill countries and mountainous regions ...
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Norway
Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and the archipelago of Svalbard also form part of Norway. Bouvet Island, located in the Subantarctic, is a dependency of Norway; it also lays claims to the Antarctic territories of Peter I Island and Queen Maud Land. The capital and largest city in Norway is Oslo. Norway has a total area of and had a population of 5,425,270 in January 2022. The country shares a long eastern border with Sweden at a length of . It is bordered by Finland and Russia to the northeast and the Skagerrak strait to the south, on the other side of which are Denmark and the United Kingdom. Norway has an extensive coastline, facing the North Atlantic Ocean and the Barents Sea. The maritime influence dominates Norway's climate, with mild lowland temperatures on the se ...
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Svalbard
Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group range from 74° to 81° north latitude, and from 10° to 35° east longitude. The largest island is Spitsbergen, followed by Nordaustlandet and . The largest settlement is Longyearbyen. The islands were first used as a base by the whalers who sailed far north in the 17th and 18th centuries, after which they were abandoned. Coal mining started at the beginning of the 20th century, and several permanent communities were established. The Svalbard Treaty of 1920 recognizes Norwegian sovereignty, and the 1925 Svalbard Act made Svalbard a full part of the Kingdom of Norway. They also established Svalbard as a free economic zone and a demilitarized zone. The Norwegian Store Norske and the Russian remain the only mining companies in place. Res ...
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Anchipteraspididae
Anchipteraspididae is an extinct family of heterostracan vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...s restricted to Late Silurian and Early Devonian strata of Arctic Canada.Elliott, D. K. "A new subfamily of the Pteraspididae (Agnatha, Heterostraci) from the upper Silurian and lower Devonian of Arctic Canada." Palaeontology 27.1 (1984): 169-197 Anchipteraspidids superficially resemble the ancestral cyathaspidiformes, cyathaspidids, but, the articulation and growth patterns of the plates clearly define them as pteraspidids. References * On the recent discovery of Pteraspidian fish in the upper Silurian rocks of North America. Edward Waller Claypole, Quart. Journ. Geol. Soc. 1885, volume 41, pages 48–64, * Revised classification of Pteraspididae with descri ...
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Benthic
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "the depths." Organisms living in this zone are called benthos and include microorganisms (e.g., bacteria and fungi) as well as larger invertebrates, such as crustaceans and polychaetes. Organisms here generally live in close relationship with the substrate and many are permanently attached to the bottom. The benthic boundary layer, which includes the bottom layer of water and the uppermost layer of sediment directly influenced by the overlying water, is an integral part of the benthic zone, as it greatly influences the biological activity that takes place there. Examples of contact soil layers include sand bottoms, rocky outcrops, coral, and bay mud. Description Oceans The benthic region of the ocean begins at the shore line (intertidal ...
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