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Parvicursorinae
Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests. '' Alvarezsaurus'', the type genus of the family, was named for the historian Gregorio Álvarez. History of study Bonaparte (1991) described the first alvarezsaurid, ''Alvarezsaurus calvoi'', from an incomplete skeleton found in Patagonia, Argentina. Bonaparte also named a family, Alvarezsauridae, to contain it. He ...
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Ondogurvel
''Ondogurvel'' () (meaning "egg lizard") is a genus of alvarezsaurid dinosaur from the Late Cretaceous (Campanian) Barun Goyot Formation in southern Mongolia. The type and only species is ''O. alifanovi'', known from a partial skeleton consisting of fragments of two last dorsal vertebrae, three anterior sacral vertebrae, right ilium, left and right pubis and ischium, articulated right tibia, fibula, metatarsals II and IV, and phalanges IV-1 and IV-2, right carpometacarpus, left and right manual phalanx II-1, right femur, left pedal phalanx II-1, and fragments of unidentified phalanges. Discovery and naming In 2022, the type species ''Ondogurvel alifanovi'' was named and described by Alexander O. Averianov and Alexey V. Lopatin. The generic name, "''Ondogurvel''" combines the Mongolian words өндөг “ondo” (), meaning egg, and гүрвэл “gurvel” (), meaning lizard. The specific name, "''alifanovi''" () honors the late Russian paleontologist Vladimir Alifanov who ...
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Qiupanykus
''Qiupanykus'' (meaning "Qiupa claw" after the Qiupa Formation) is a genus of alvarezsaurid coelurosaur theropod from the Late Cretaceous Qiupa Formation of southern China. Fossil eggs believed to be those of an oviraptorid found in association with the holotype specimen indicate that both ''Qiupanykus'' and other alvarezsaurids may have been specialist egg eaters that used their robust thumb claws to crack open eggshells. Phylogeny Lü Junchang ''et al.'' (2018) recovered ''Qiupanykus'' as a member of the Alvarezsauridae, falling closer to ''Parvicursor'' than to ''Patagonykus''. Although not formally assigned to the clade Parvicursorinae Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly ... in the description, this position would make it a member of Parvicursorinae ''sensu'' Xu '' ...
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Bonapartenykus
''Bonapartenykus'' (meaning " José F. Bonaparte's claw") is a monospecific genus of alvarezsauroid dinosaur from Argentina that lived during the Late Cretaceous (Campanian-Maastrichtian) in what is now the upper Allen Formation of the Río Negro Province. The type and only species, ''Bonapartenykus ultimus'', is known from a nearly articulated but partial skeleton that was found in close association to two incomplete eggs and several clusters of eggshells belonging to the oogenus '' Arriagadoolithus''. ''Bonapartenykus'' was named in 2012 by Federico L. Agnolin, Jaime E. Powell, Fernando E. Novas and Martin Kundrát. ''Bonapartenykus'' has an estimated length of and weight of , making it the largest member of the clade Alvarezsauroidea. Discovery and Naming A partial skeleton of a theropod with eggs was collected in a surface of approximately 30 m2 in fluvial sandstones of the upper Allen Formation in northwestern Patagonia, Argentina. The locality has also produced specimen ...
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Dzharaonyx
''Dzharaonyx'' (meaning "Dzharakuduk claw", named after the type locality) is a genus of alvarezsaurid theropod dinosaur from the Late Cretaceous Bissekty Formation of Uzbekistan. The type species is ''Dzharaonyx eski''; ''eski'' being an Uzbek word for "old". Description ''Dzharaonyx'' is known from a series of disassociated but well-preserved postcranial material. The form of the humerus is intermediate between that of '' Patagonykus'' and ''Mononykus''. Classification Phylogenetic analysis places ''Dzharaonyx'' in a polytomy including other Asian members of Parvicursorinae. This makes it the oldest parvicursorine known to date. Paleoenvironment ''Dzharaonyx'' lived in the Bissekty Formation, which has been intensively studied in recent years. It represents a brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt ...
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Ceratonykus
''Ceratonykus'' (meaning "horned claw") is a monospecific genus of alvarezsaurid dinosaur from Mongolia that lived during the Late Cretaceous (late Campanian stage, 72.1 Ma) in what is now the Barun Goyot Formation. The type and only species, ''Ceratonykus oculatus'', is known from a fragmentary skeleton, including an incomplete skull, of an adult individual. It was named and described in 2009 by Vladimir Alifanov and Rinchen Barsbold. Its describers questioned the traditional placement of alvarezsaurs in Theropoda, instead suggesting they were ornithischians, but this has not been accepted since. ''Ceratonykus'' has an estimated length of 75 centimetres (2.5 feet) and weight of 760 grams (1.68 lbs). It has been considered as a possible junior synonym of '' Parvicursor''. Discovery and naming In 2003, a partial skeleton of an alvarezsaurid was discovered in the Barun Goyot Formation of the Khermiin Tsav locality, Mongolia by the Paleontological Center of the Mongolian Acade ...
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Patagonykus
''Patagonykus'' (meaning "Patagonian claw") is a genus of theropod dinosaur from the Upper Cretaceous of Argentina. This alvarezsauroid was discovered in exposures of the Portezuelo Formation (Turonian- Coniacian) of the Rio Neuquén Subgroup in the Neuquén Basin, Neuquen Province of Patagonia, Argentina. The holotype consists of an incomplete but well-preserved skeleton, lacking a skull, but including many vertebrae, the coracoids, a partial forelimb, pelvic girdle, and hindlimbs. ''Patagonykus'' has been classed with the Alvarezsauridae, a family which includes such taxa as the Mongolian ''Mononykus'' and the Argentinian ''Alvarezsaurus''. In 2010 Gregory S. Paul estimated its length at 1 meter (3.3 ft) and its weight at 3.5 kg (7.7 lbs). Classification Agnolin ''et al.'' (2012) originally placed ''Patagonykus'' within Alvarezsauridae, within the clade Patagonykinae as sister taxon to '' Bonapartenykus''. Makovicky, Apesteguía and Gianechini (2012) found it to be in a po ...
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Khulsanurus
''Khulsanurus'' (meaning "tail from Khulsan") is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous Barungoyot Formation of the Khulsan Locality in the Gobi Desert region of Mongolia. The type and only species is ''Khulsanurus magnificus''. Description ''Khulsanurus'' is known from a holotype containing partial vertebral series of the neck and tail, scapulocoracoids, a right humerus, a possible left humerus, and a left pubis. Distinguishing autapomorphies of this specimen include pleurocoel-, carotid process-, and epipophysis-less cervical vertebrae, prominent infrapostzygapophyseal fossa of the caudal vertebrae which also bear anteriorly pointing prezygapophyses. Classification Averianov & Lopatin (2021) placed ''Khulsanurus'' in a polytomy with '' Albinykus'', ''Ceratonykus'', '' Linhenykus'', '' Parvicursor'', '' Xixianykus'', ''Qiupanykus'', ''Nemegtonykus'', PIN 5838/1 (=''Ondogurvel'') and a clade containing '' Shuvuuia'' and ''Mononykus'' in ...
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Parvicursor
''Parvicursor'' (meaning "small runner") is a genus of tiny maniraptoran dinosaur with long slender legs for fast running. At only about from snout to end of tail, and in weight, it was initially seen as one of the smallest non-avian dinosaurs known from an adult specimen.Which was the smallest dinosaur?
Royal Tyrrell Museum. Last accessed 2008-05-23.
However, in 2022 its was concluded to represent a juvenile individual. Like other members of the

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Linhenykus
''Linhenykus'' is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous of Inner Mongolia, China. It is the most basal known member of the Parvicursorinae. The genus gets its name from Linhe, a city near the site where the fossil was first found and Greek ''nykus'', "claw". The specific name is derived from Greek ''monos'', "single", and ''daktylos'', "finger", a reference to the fact that it is the only known non-avian dinosaur to have had but a single digit. Description ''Linhenykus'' was a small dinosaur, measuring long and weighing . Its femur length is . Alvarezsauroids are known for their short forelimbs, each with a single greatly enlarged second digit. Although alvarezsaurids were once thought to have only a single digit on each forelimb, more recent evidence has shown that most species have reduced third and fourth digits. ''Linhenykus'' is the first known alvarezsaurid to have only a single, second digit. Although a reduced third metacarpal ...
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Trierarchuncus
''Trierarchuncus'' (meaning "captain hook," after its single-clawed hands) is a monotypic genus of alvarezsaurid theropod which includes a single species, ''Trierarchuncus prairiensis'', which is known from fossils found in deposits of the Hell Creek Formation in Montana. It is the youngest known alvarezsaurid and one of the last non-avian dinosaurs, going extinct during the Cretaceous–Paleogene extinction event, which occurred approximately 66 million years ago. Discovery and naming The first remains ware discovered in Montana in 1980 and it was informally known as the "Hell Creek alvarezsaur".Hutchinson and Chiappe, 1998. The first known alvarezsaurid (Theropoda: Aves) from North America. ''Journal of Vertebrate Paleontology''. 18(3), 447–450. The then unnamed species was not mentioned again until it was mentioned briefly in the 2018 Society of Vertebrate Paleontology abstract book. The species and genus were scientifically described by Denver Fowler and colleagues in 2020 b ...
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Cyril Walker (ornithologist)
Cyril Alexander Walker (8 February 1939 – 6 May 2009) was a British palaeontologist, curator of fossil birds in the Natural History Museum. He was also interested in fossil turtles. book, '' Smithsonian Handbook of Fossils''. He has also contributed to many other books, including ''Garden Birds'', ''Field Guide to British Birds'', ''Birds of the World'', ''Nature Notebooks'', and others. An unidentified moa bone of unknown origin and locality, donated by Dr. C. Walker to ornithologist Zlatozar Boev in 1986, was later identified as the little bush moa (''Anomalopteryx didiformis'').1. Boev, Z. 2018. A specimen of little bush moa Anomalopteryx didiformis (Owen, 1844), Emeidae Bonaparte, 1854 from the National Museum of Natural History, Sofia. – Historia naturalis bulgarica, 32: 3-5. It is the only specimen of Dinornithiformes in the collections of the National Museum of Natural History, Bulgarian Academy of Sciences in Sofia. References British palaeontologists Briti ...
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