Parvicursor Remotus
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Parvicursor Remotus
''Parvicursor'' (meaning "small runner") is a genus of tiny maniraptoran dinosaur with long slender legs for fast running. At only about from snout to end of tail, and in weight, it was initially seen as one of the smallest non-avian dinosaurs known from an adult specimen.Which was the smallest dinosaur?
Royal Tyrrell Museum. Last accessed 2008-05-23.
However, in 2022 its was concluded to represent a juvenile individual. Like other members of the

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Nest
A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic material such as twigs, grass, and leaves, or may be a simple depression in the ground, or a hole in a rock, tree, or building. Human-made materials, such as string, plastic, cloth, or paper, may also be used. Nests can be found in all types of habitat. Nest building is driven by a biological urge known as the nesting instinct in birds and mammals. Generally each species has a distinctive style of nest. Nest complexity is roughly correlated with the level of parental care by adults. Nest building is considered a key adaptive advantage among birds, and they exhibit the most variation in their nests ranging from simple holes in the ground to elaborate communal nests hosting hundreds of individuals. Nests of prairie dogs and several social insec ...
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Campanian
The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian spans the time from 83.6 (± 0.2) to 72.1 (± 0.2) million years ago. It is preceded by the Santonian and it is followed by the Maastrichtian. The Campanian was an age when a worldwide sea level rise covered many coastal areas. The morphology of some of these areas has been preserved: it is an unconformity beneath a cover of marine sedimentary rocks. Etymology The Campanian was introduced in scientific literature by Henri Coquand in 1857. It is named after the French village of Champagne in the department of Charente-Maritime. The original type locality was a series of outcrop near the village of Aubeterre-sur-Dronne in the same region. Definition The base of the Campanian Stage is defined as a place in the stratigraphic column wher ...
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Alvarezsaurids
Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests. ''Alvarezsaurus'', the type genus of the family, was named for the historian Gregorio Álvarez. History of study Bonaparte (1991) described the first alvarezsaurid, ''Alvarezsaurus calvoi'', from an incomplete skeleton found in Patagonia, Argentina. Bonaparte also named a family, Alvarezsauridae, to contain it. He a ...
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Ceratonykus
''Ceratonykus'' (meaning "horned claw") is a monospecific genus of alvarezsaurid dinosaur from Mongolia that lived during the Late Cretaceous (late Campanian stage, 72.1 Ma) in what is now the Barun Goyot Formation. The type and only species, ''Ceratonykus oculatus'', is known from a fragmentary skeleton, including an incomplete skull, of an adult individual. It was named and described in 2009 by Vladimir Alifanov and Rinchen Barsbold. Its describers questioned the traditional placement of alvarezsaurs in Theropoda, instead suggesting they were ornithischians, but this has not been accepted since. ''Ceratonykus'' has an estimated length of 75 centimetres (2.5 feet) and weight of 760 grams (1.68 lbs). It has been considered as a possible junior synonym of ''Parvicursor''. Discovery and naming In 2003, a partial skeleton of an alvarezsaurid was discovered in the Barun Goyot Formation of the Khermiin Tsav locality, Mongolia by the Paleontological Center of the Mongolian Academy of ...
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Linhenykus
''Linhenykus'' is an extinct genus of alvarezsaurid theropod dinosaur from the Late Cretaceous of Inner Mongolia, China. It is the most basal known member of the Parvicursorinae. The genus gets its name from Linhe, a city near the site where the fossil was first found and Greek ''nykus'', "claw". The specific name is derived from Greek ''monos'', "single", and ''daktylos'', "finger", a reference to the fact that it is the only known non-avian dinosaur to have had but a single digit. Description ''Linhenykus'' was a small dinosaur, measuring long and weighing . Its femur length is . Alvarezsauroids are known for their short forelimbs, each with a single greatly enlarged second digit. Although alvarezsaurids were once thought to have only a single digit on each forelimb, more recent evidence has shown that most species have reduced third and fourth digits. ''Linhenykus'' is the first known alvarezsaurid to have only a single, second digit. Although a reduced third metacarpal ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Phil Currie
Philip John Currie (born March 13, 1949) is a Canadian palaeontologist and museum curator who helped found the Royal Tyrrell Museum of Palaeontology in Drumheller, Alberta and is now a professor at the University of Alberta in Edmonton. In the 1980s, he became the director of the Canada-China Dinosaur Project, the first cooperative palaeontological partnering between China and the West since the Central Asiatic Expeditions in the 1920s, and helped describe some of the first feathered dinosaurs. He is one of the primary editors of the influential ''Encyclopedia of Dinosaurs'', and his areas of expertise include theropods (especially Tyrannosauridae), the origin of birds, and dinosaurian migration patterns and herding behavior. He was one of the models for palaeontologist Alan Grant in the film ''Jurassic Park''. Biography Currie received his Bachelor of Science degree from the University of Toronto in 1972, a Master of Science degree from McGill University in 1975, and a Doc ...
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Parvicursorinae
Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests. ''Alvarezsaurus'', the type genus of the family, was named for the historian Gregorio Álvarez. History of study Bonaparte (1991) described the first alvarezsaurid, ''Alvarezsaurus calvoi'', from an incomplete skeleton found in Patagonia, Argentina. Bonaparte also named a family, Alvarezsauridae, to contain it. He a ...
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Mononykus
''Mononykus'' ( , sometimes ; meaning "one claw") is a genus of alvarezsaurid dinosaur that lived during the Late Cretaceous in what is now Asia on the Nemegt Formation, about 70 million years ago. ''Mononykus'' was a very small theropod, estimated around in length with a weight of . As in ''Shuvuuia'', ''Mononykus'' likely developed a shaggy feathering. It was lightly built with long, thin legs and highly reduced and specialized forelimbs that were likely used for foraging termite mounds or other insect colonies. History of discovery ''Mononykus'' is represented by a single holotype specimen, catalog number MPC-D 107/6 (formerly IGM 107/6). It was collected in 1987 from the Bügiin Tsav locality of the Nemegt Formation, Gobi Desert. This specimen consists of a partial skeleton lacking a tail, and only small fragments of skull bones, including a complete braincase. ''Mononykus'' was originally named ''Mononychus'' in 1993, but later that year, it was renamed because the origin ...
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Shuvuuia
''Shuvuuia'' is a genus of bird-like theropod dinosaur from the late Cretaceous period of Mongolia. It is a member of the family Alvarezsauridae, small coelurosaurian dinosaurs which are characterized by short but powerful forelimbs specialized for digging. The type (and only known) species is ''Shuvuuia deserti'', or "desert bird".Chiappe, L.M., Norell, M. A., and Clark, J. M. (1998). "The skull of a relative of the stem-group bird ''Mononykus''." ''Nature'', 392(6673): 275-278. The name ''Shuvuuia'' is derived from the Mongolian word ''shuvuu'' (шувуу) meaning "bird". Description ''Shuvuuia'' was a small and lightly built animal. At in length and in body mass, it is one of the smallest known non-avian dinosaurs. The skull is lightly built with long and slender jaws and minute teeth. ''Shuvuuia'' is unique among non-avian theropods in the skull's ability to perform prokinesis, that is, it could flex its upper jaw independently of its braincase. The hindlimbs of ''Shuvu ...
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Pelvis
The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The pelvic region of the trunk includes the bony pelvis, the pelvic cavity (the space enclosed by the bony pelvis), the pelvic floor, below the pelvic cavity, and the perineum, below the pelvic floor. The pelvic skeleton is formed in the area of the back, by the sacrum and the coccyx and anteriorly and to the left and right sides, by a pair of hip bones. The two hip bones connect the spine with the lower limbs. They are attached to the sacrum posteriorly, connected to each other anteriorly, and joined with the two femurs at the hip joints. The gap enclosed by the bony pelvis, called the pelvic cavity, is the section of the body underneath the abdomen and mainly consists of the reproductive organs (sex organs) and the rectum, while the pelvic f ...
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Mongolia
Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, making it the world's most sparsely populated sovereign nation. Mongolia is the world's largest landlocked country that does not border a closed sea, and much of its area is covered by grassy steppe, with mountains to the north and west and the Gobi Desert to the south. Ulaanbaatar, the capital and largest city, is home to roughly half of the country's population. The territory of modern-day Mongolia has been ruled by various nomadic empires, including the Xiongnu, the Xianbei, the Rouran, the First Turkic Khaganate, and others. In 1206, Genghis Khan founded the Mongol Empire, which became the largest contiguous land empire in history. His grandson Kublai Khan conquered China proper and established the Yuan dynasty. After the co ...
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