Pachycephalosauria Jmallon
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Pachycephalosauria Jmallon
Pachycephalosauria (; from Greek παχυκεφαλόσαυρος for 'thick headed lizards') is a clade of ornithischian dinosaurs. Along with Ceratopsia, it makes up the clade Marginocephalia. With the exception of two species, most pachycephalosaurs lived during the Late Cretaceous Period, dating between about 85.8 and 65.5 million years ago. They are exclusive to the Northern Hemisphere, all of them being found in North America and Asia. They were all bipedal, herbivorous/omnivorous animals with thick skulls. Skulls can be domed, flat, or wedge-shaped depending on the species, and are all heavily ossified. The domes were often surrounded by nodes and/or spikes. Partial skeletons have been found of several pachycephalosaur species, but to date no complete skeletons have been discovered. Often isolated skull fragments are the only bones that are found. Candidates for the earliest known pachycephalosaur include ''Ferganocephale adenticulatum'' from Middle Jurassic Period strata ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia an ...
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Foraminacephale
''Foraminacephale'' (meaning "foramina head") is a genus of pachycephalosaurid dinosaur from Late Cretaceous (Campanian stage) deposits of Canada. Description ''Foraminacephale'', as a pachycephalosaurid, was a small, bipedal herbivore with a thickened dome on its skull. In 2016, Gregory S. Paul estimated its length at , its weight at . In ''Foraminacephale'', the top surface of the dome is punctuated by many small pits, the eponymous foramina; the dome itself consists of a large, central lobe with a sloped frontal half, and two smaller lateral lobes at the front. The squamosal bone forms a tall bar of completely smooth bone underneath the dome, save for six bony nodes that line the bottom edge of the dome and an additional "corner" node just below. These features differentiate ''Foraminacephale'' from all other pachycephalosaurids. Unlike ''Stegoceras'', '' Hanssuesia'', and '' Colepiocephale'', the parietal bone of ''Foraminacephale'' (which constitutes the back part of the ...
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Marginocephalia
Marginocephalia (/mär′jə-nō-sə-făl′ē-ən/ Latin: margin-head) is a clade of ornithischian dinosaurs that is characterized by a bony shelf or margin at the back of the skull. These fringes were likely used for display. There are two clades included in Marginocephalia: the thick-skulled Pachycephalosauria and the horned Ceratopsia. All members of Marginocephalia were primarily herbivores. They basally used gastroliths to aid in digestion of tough plant matter until they convergently evolved tooth batteries in Neoceratopsia (or "new Ceratopsia") and Pachycephalosauria. Marginocephalia first evolved in the Jurassic Period and became more common in the Cretaceous. They are basally small facultative quadrupeds while derived members of the group are large obligate quadrupeds. Primitive marginocephalians are found in Asia, but the group migrated upwards into North America. Pachycephalosaurs, or "thick-headed reptiles", have primitive features that include basally small sized bo ...
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Ceratopsia
Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassic. The earliest known ceratopsian, '' Yinlong downsi'', lived between 161.2 and 155.7 million years ago.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> The last ceratopsian species, '' Triceratops prorsus'', became extinct during the Cretaceous–Paleogene extinction event, . ''Triceratops'' is by far the best-known ceratopsian to the general public. It is traditional for ceratopsian genus names to end in "''-ceratops''", although this is not always the case. One of the first named genera was '' Ceratops'' itself, which lent its name to the group, although it is considered a ''nomen dubium'' today as its fossil remains have no distinguishing ch ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, ar ...
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Ornithischian
Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek stem ' (), meaning "of a bird", and ' (), plural ', meaning "hip joint". However, birds are only distantly related to this group as birds are theropod dinosaurs. Ornithischians with well known anatomical adaptations include the ceratopsians or "horn-faced" dinosaurs (e.g. ''Triceratops''), the pachycephalosaurs or "thick-headed" dinosaurs, the armored dinosaurs ( Thyreophora) such as stegosaurs and ankylosaurs, and the ornithopods. There is strong evidence that certain groups of ornithischians lived in herds, often segregated by age group, with juveniles forming their own flocks separate from adults. Some were at least partially covered in filamentous (hair- or feather- like) pelts, and there is much debate over whether these filaments foun ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between org ...
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Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic period (), and the Classical period (). Ancient Greek was the language of Homer and of fifth-century Athenian historians, playwrights, and philosophers. It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article primarily contains information about the Epic and Classical periods of the language. From the Hellenistic period (), Ancient Greek was followed by Koine Greek, which is regarded as a separate historical stage, although its earliest form closely resembles Attic Greek and its latest form approaches Medieval Greek. There were several regional dialects of Ancient Greek, of which Attic Greek developed into Koi ...
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Oxford University Press
Oxford University Press (OUP) is the university press of the University of Oxford. It is the largest university press in the world, and its printing history dates back to the 1480s. Having been officially granted the legal right to print books by decree in 1586, it is the second oldest university press after Cambridge University Press. It is a department of the University of Oxford and is governed by a group of 15 academics known as the Delegates of the Press, who are appointed by the vice-chancellor of the University of Oxford. The Delegates of the Press are led by the Secretary to the Delegates, who serves as OUP's chief executive and as its major representative on other university bodies. Oxford University Press has had a similar governance structure since the 17th century. The press is located on Walton Street, Oxford, opposite Somerville College, in the inner suburb of Jericho. For the last 500 years, OUP has primarily focused on the publication of pedagogical texts an ...
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Lexico
Lexico was a dictionary website that provided a collection of English and Spanish dictionaries produced by Oxford University Press (OUP), the publishing house of the University of Oxford. While the dictionary content on Lexico came from OUP, this website was operated by Dictionary.com, whose eponymous website hosts dictionaries by other publishers such as Random House. The website was closed and redirected to Dictionary.com on 26 August 2022. Before the Lexico site was launched, the '' Oxford Dictionary of English'' and '' New Oxford American Dictionary'' were hosted by OUP's own website Oxford Dictionaries Online (ODO), later known as Oxford Living Dictionaries. The dictionaries' definitions have also appeared in Google definition search and the Dictionary application on macOS, among others, licensed through the Oxford Dictionaries API. History In the 2000s, OUP allowed access to content of the ''Compact Oxford English Dictionary of Current English'' on a website called As ...
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Alaskacephale
''Alaskacephale'' was a genus of pachycephalosaurid dinosaur from the Prince Creek Formation, that lived in the early Maastrichtian stages of the Late Cretaceous (around 71 to 69 million years ago). Discovery ''Alaskacephale'' was named by Robert Sullivan in 2006. The genus name refers to Alaska, where the holotype was discovered, combined with the Greek ''kephale'', meaning head. The species name, ''gangloffi'', honors paleontologist Roland Gangloff. The only known specimen of ''A. gangloffi'' is the holotype UAM AK-493-V-001, found in 1999, a nearly complete left squamosal with a characteristic array of polygonal nodes. The dimensions of this bone suggest that ''A. gangloffi'' was about half the size of '' Pachycephalosaurus wyomingensis'' or three quarters the size of '' Prenocephale'', and about the same size as '' Sphaerotholus edmontonensis'' and ''Foraminacephale''. The specimen was previously described by Gangloff ''et al.'' (2005) as an unnamed pachycephalosaurid, pos ...
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Tylocephale
''Tylocephale'' (meaning "swollen head", from the Greek ''τυλη'' meaning 'callus' or 'hard swelling' and ''κεφαλη'' meaning 'head') is a genus of pachycephalosaurid dinosaur from the Late Cretaceous period. It was a herbivorous dinosaur, reaching in length and in body mass. It had the tallest dome of any known pachycephalosaur. ''Tylocephale'' lived during the Campanian stage, around 74 million years ago. It was discovered in the Khulsan locality of the Barun Goyot Formation, Mongolia sometime between 1965 and 1971. The type species is ''T. gilmorei'', described by Teresa Maryańska and Osmólska in 1974 based on a partial skull (specimen ZPAL MgD-I/105). Pachycephalosaurids evolved in Asia and then migrated into North America, thus it is likely that ''Tylocephale'' migrated back into Asia. It is closely related to '' Prenocephale''. See also * Timeline of pachycephalosaur research This timeline of pachycephalosaur research is a chronological listing of e ...
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