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Pytine
''Pytine'' is an extinct genus of asaphida, asaphid trilobites. Species lived during the later part of the Arenig, Arenig stage of the Ordovician Period, approximately 478 to 471 million years ago. Various species are found in the Svalbard, Valhallfonna Formation, Olenidsletta, Member, of Spitzbergen, Norway, the ''Megistaspis (Paramegistaspis) planilimbata'' Zone of the '''Shumardia'' Shale' of Sweden, Jujuy Province, Argentina, early Arenig-aged strata of Jiangxi province, China, and Darriwilian-aged strata in Western Hunan province, China. The type species, ''P. graia'', has seven thorax segments, and lacks the rapier-like glabellar spine, that occurs in many other raphiophorids. The Chinese species, by contrast, have only six thoracic segments. So far, only the type species, and one of the Chinese species, ''P. laevigata'' (synonym= ''Jiuxiella laevigata''), are known from complete specimens. Species and synonymy Richard Fortey erected the genus in 1975 upon describing ''P ...
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Raphiophoridae
Raphiophoridae is a family of small to average-sized trilobites that first occurred at the start of the Ordovician and became extinct at the end of the Middle Silurian. Anatomy All raphiophorids are blind, with headshields (or cephalons) that are triangular to subcircular, and many carry long, trailing genal spines, a forward-directed rapier-like spine on the central raised area (or glabella), or both, with the glabella often inflated and the natural fracture lines (or sutures) of the cephalon coinciding with its margin. The thorax typically has five to seven segments (except for the genera ''Taklamakania'', ''Pseudampyxina'', ''Nanshanaspis'', and ''Kongqiangheia'', which have only 3). As mentioned before, all raphiophorids are blind. Many, if not most, genera have no eyes whatsoever, though a few, such as '' Lehnertia'', have vestigial tubercles that correspond to the compound eyes of their ancestors. Distribution Raphiophoridae currently includes two officially recog ...
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Arenig
In geology, the Arenig (or Arenigian) is a time interval during the Ordovician period and also the suite of rocks which were deposited during this interval. History The term was first used by Adam Sedgwick in 1847 with reference to the "Arenig Ashes and Porphyries" in the neighbourhood of Arenig Fawr, in Merioneth, North Wales. The rock-succession in the Arenig district has been recognized by W. G. Fearnsides (β€œOn the Geology of Arenig Fawr and Moel Llanfnant", Q.J.G.S. vol. lxi., 1905, pp. 608–640, with maps). The above succession is divisible into: # A lower series of gritty and calcareous sediments, the "Arenig Series" as it is now understood; # A middle series, mainly volcanic, with shale, the "Llandeilo Series"; and # The shale and limestones of the Bala or Caradoc Stage. It was to the middle series (2) that Sedgwick first applied the term "Arenig". In the typical region and in North Wales generally the Arenig series appears to be unconformable upon the Cambri ...
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Richard Fortey
Richard Alan Fortey FRS FRSL (born 15 February 1946 in London) is a British palaeontologist, natural historian, writer and television presenter, who served as president of the Geological Society of London for its bicentennial year of 2007. Early life and education Fortey was educated at Ealing Grammar School for Boys and King's College, Cambridge, where he read Natural Sciences specialising in geology. He received a PhD and DSc from the University of Cambridge. Career Fortey has had a long career as a palaeontologist at the Natural History Museum in London; his research interests include above all, trilobites: at the age of 14, he discovered his first trilobite, sparking a passionate interest that later became a career. He has named numerous trilobite species and still continues his research despite having retired from the Museum. He studies trilobites and graptolites, especially those from the Ordovician and their systematics, evolution and modes of life; he is also involved ...
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Ampyx (trilobite)
''Ampyx'' is an Ordovician-Silurian genus of Asaphid trilobites of the family Raphiophoridae. Species of ''Ampyx'' are characterized by three extended spines on the head-shield, one spine derived from each free cheek, and one spine emanating from the glabellum. Species include ''Ampyx linleyensis'' ( Lanvirn-Caradoc series). Species of ''Ampyx'' grew to an average length of . Collective behaviour Fossils of the trilobite ''Ampyx priscus'', dating back about 480 million years ago, have been recently described as clustered in lines along the ocean floor. The animals were all mature adults, and were all facing the same direction as though they had formed a conga line or a peloton. It has been suggested they line up in this manner to migrate, much as spiny lobsters migrate in single-file queues. Or perhaps they are getting together for mating. The findings suggest animal collective behaviour has very early evolutionary origins.Vannier J, Vidal M, Marchant R, El Hariri K, Kourais ...
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Extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. More than 99% of all species that ever lived on Earth, amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryote globally, and possibly many times more if microorganisms, like bacteria, are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, dodos, m ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Asaphida
Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies (Anomocaroidea, Asaphoidea, Cyclopygoidea, Dikelocephaloidea, Remopleuridoidea and Trinucleioidea), but no suborders. Asaphids comprise some 20% of described fossil trilobites. In 2020, the superfamily Trinucleoidea was proposed to be raised to an order ( Trinucleida) and removed from Asaphida. Morphology The Asaphids generally have cephalon (head) and pygidium (tail) parts similar in size, and most species have a prominent median ventral suture. Heads are often flat, and carapace furrows in the head area are often faint or not visible. Thoracic segments typically number 5 - 12, though some species have as few as two and some as many as 30. They also generally have a wide doublure, or rim, that surrounds the cephalon. This causes some specimens to be described as having a char ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The stu ...
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Ordovician Period
The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Geological Congress. Life continued to flourish during the Ordovician as it did in the earlier Cambrian P ...
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Ancient Greek
Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic period (), and the Classical period (). Ancient Greek was the language of Homer and of fifth-century Athenian historians, playwrights, and philosophers. It has contributed many words to English vocabulary and has been a standard subject of study in educational institutions of the Western world since the Renaissance. This article primarily contains information about the Epic and Classical periods of the language. From the Hellenistic period (), Ancient Greek was followed by Koine Greek, which is regarded as a separate historical stage, although its earliest form closely resembles Attic Greek and its latest form approaches Medieval Greek. There were several regional dialects of Ancient Greek, of which Attic Greek developed into Koine. Dia ...
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Asaphida Genera
Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies (Anomocaroidea, Asaphoidea, Cyclopygoidea, Dikelocephaloidea, Remopleuridoidea and Trinucleioidea), but no suborders. Asaphids comprise some 20% of described fossil trilobites. In 2020, the superfamily Trinucleoidea was proposed to be raised to an order ( Trinucleida) and removed from Asaphida. Morphology The Asaphids generally have cephalon (head) and pygidium (tail) parts similar in size, and most species have a prominent median ventral suture. Heads are often flat, and carapace furrows in the head area are often faint or not visible. Thoracic segments typically number 5 - 12, though some species have as few as two and some as many as 30. They also generally have a wide doublure, or rim, that surrounds the cephalon. This causes some specimens to be described as having a charact ...
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