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Provora
Provora is a proposed supergroup of eukaryotes made up of predatory microbes, "devouring voracious protists". It was reported that ten strains were isolated and cultured in 2022. They are predators of other microorganisms. Their discovery was very delayed, compared to other microorganisms in their environments, due to their rarity. Their 18S is very different from that of other eukaryotes, thus they were taxonomically placed in a separate supergroup. Phylogeny External relationships The supergroup Provora is composed of eukaryotic strains that form an ancient lineage within the clade Diaphoretickes. Phylogenetic analyses have recovered the following cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... shape: Internal relationships The phylogenetic relationships between th ...
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Supergroup (biology)
A supergroup, in evolutionary biology, is a large group of organisms that share one common ancestor and have important defining characteristics. It is an informal, mostly arbitrary rank in biological taxonomy that is often greater than phylum or kingdom, although some supergroups are also treated as phyla. Eukaryotic supergroups Since the decade of 2000's, the eukaryotic tree of life (abbreviated as eToL) has been divided into 5–8 major groupings called 'supergroups'. These groupings were established after the idea that only monophyletic groups should be accepted as ranks, as an alternative to the use of paraphyletic kingdom Protista. In the early days of the eToL six traditional supergroups were considered: Amoebozoa, Opisthokonta, "Excavata", Archaeplastida, "Chromalveolata" and Rhizaria. Since then, the eToL has been rearranged profoundly, and most of these groups were found as paraphyletic or lacked defining morphological characteristics that unite their members, which mak ...
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Eukaryotes
Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard archaea. This implies that there are only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as flagellated phagotrophs. Their name comes from the Greek εὖ (''eu'', "well" or "good") and κάρυον (''karyon'', "nut" or "kernel"). Eu ...
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Ancoracysta Twista
''Ancoracysta twista'' is a eukaryotic microorganism. It is a predatory protist that appears to be sister to Haptista.Janouškovec J, Tikhonenkov DV, Burki F, Howe AT, Rohwer FL, Mylnikov AP, Keeling PJ. "A New Lineage of Eukaryotes Illuminates Early Mitochondrial Genome Reduction", ''Current Biology'' 2017, Description ''Ancoracysta twista'' was first described in November 2017 in ''Current Biology''. It was found in a sample collected from the surface of a tropical aquarium brain coral. It actively feeds on '' Procryptobia sorokini'', probably immobilising its prey through discharging a previously unknown type of extrusome named an ancoracyst. Genetic analysis shows that it is not closely related to any known lineage, but it may be most closely related to a grouping of haptophytes and centrohelids (Haptista). It is notable for having a gene-rich mitochondrial genome, the largest known outside the jakobids or '' Diphylleia rotans''. Uniquely, it appears to contain both the nu ...
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Haptista
Haptista is a proposed group of protists made up of centrohelids and haptophytes. Phylogenomic studies indicate that Haptista, together with ''Ancoracysta twista'', forms a sister clade to the SAR+Telonemia supergroup, but it may also be sister to the Cryptista (+Archaeplastida). It is thus one of the earliest diverging Diaphoretickes. Taxonomy Based on studies done by Cavalier-Smith, Chao & Lewis 2015 and Ruggiero et al. 2015. * Subphylum Centroheliozoa Cushman & Jarvis 1929 sensu Durrschmidt & Patterson 1987 eliozoa Haeckel 1862 stat. n. Margulis 1974 em. Cavalier-Smith 2003** Class Centrohelea Kuhn 1926 stat. n. Cavalier-Smith 1993 entroplastiales; Centrohelina Hartmann 1913; Centroplasthelida Febvre-Chevalier 1984* Subphylum Haptophytina Cavalier-Smith 2015 (Haptophyta Hibberd 1976 sensu Ruggerio et al. 2015) ** Clade Rappemonada Kim et al. 2011 *** Class Rappephyceae Rappephyceae, or Rappemonads, are a small family of protists first described in 2011, of uncertain phylog ...
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Archaeplastida
The Archaeplastida (or kingdom Plantae ''sensu lato'' "in a broad sense"; pronounced /ɑːrkɪ'plastɪdə/) are a major group of eukaryotes, comprising the photoautotrophic red algae (Rhodophyta), green algae, land plants, and the minor group glaucophytes. It also includes the non-photosynthetic lineage Rhodelphidia, a predatorial (eukaryotrophic) flagellate that is sister to the Rhodophyta, and probably the microscopic picozoans. The Archaeplastida have chloroplasts that are surrounded by two membranes, suggesting that they were acquired directly through a single endosymbiosis event by feeding on a cyanobacterium. All other groups which have chloroplasts, besides the amoeboid genus ''Paulinella'', have chloroplasts surrounded by three or four membranes, suggesting they were acquired secondarily from red or green algae. Unlike red and green algae, glaucophytes have never been involved in secondary endosymbiosis events. The cells of the Archaeplastida typically lack centriol ...
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