Proterorhinus Nasalis
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Proterorhinus Nasalis
The eastern tubenose goby (''Proterorhinus nasalis'') is a species of gobiid fish native to fresh and brackish waters of the basins of the Sea of Azov and the Caspian Sea and has invaded the upper reaches of the Volga River from its native occurrence in the delta. This species prefers slow flowing rivers or still waters with plentiful rocks or vegetation. It can reach a length of SL. It is probably the same species as that recently treated as ''Proterorhinus semipellucidus ''Proterorhinus semipellucidus '' is a species of gobiid fish, a tubenose goby originally described from the Gharasu River near Gorgan Bay of the Caspian Sea in Iran. Following the systematic decomposition of the tubenose gobies it was sugge ...'' (or ''P.'' cf. ''semipellucidus'').Brian W. CoadFreshwater Fishes of Iran(accessed 22 Feb 2015) References

Proterorhinus, Eastern tubenose goby Fish of the Caspian Sea Fish of Western Asia Fish described in 1963 Endemic fauna of the Caspian Sea T ...
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Jörg Freyhof
Jörg Arthur Freyhof (born 4 November 1964 in Ludwigshafen) is a German ichthyologist specializing on Old World cypriniform fishes.Dr. Jörg Freyhof
Leibniz Institute of Freshwater Ecology and Inland Fisheries
Freyhof has worked at the Alexander Koenig Research Museum, Bonn, and since 2000 he has been employed at the
Leibniz Institute of Freshwater Ecology and Inland Fisheries Gottfried Wilhelm (von) Leibniz . ( – 14 November 1716) was a German polymath active as a mathematician, philosopher, scientis ...
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Volga River
The Volga (; russian: Во́лга, a=Ru-Волга.ogg, p=ˈvoɫɡə) is the List of rivers of Europe#Rivers of Europe by length, longest river in Europe. Situated in Russia, it flows through Central Russia to Southern Russia and into the Caspian Sea. The Volga has a length of , and a catchment area of «Река Волга»
, Russian State Water Registry
which is more than twice the size of Ukraine. It is also Europe's largest river in terms of average discharge (hydrology), discharge at delta – between and – and of drainage basin. It is widely regarded as the Rivers in Russia, national river of Russia. The hypothetical old Russian state, the Rus' Khaganate, arose along the Volga . Historically, the river served as an important meeting place of various Eurasian civilizations. The river flows in Russia through forests, Fo ...
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Fish Described In 1963
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of living fish species are ray-finned fish, belonging to the class Actinopterygii, with around 99% of those being teleosts. The earliest organisms that can be classified as fish were soft-bodied chordates that first appeared during the Cambrian period. Although they lacked a true spine, they possessed notochords which allowed them to be more agile than their invertebrate counterparts. Fish would continue to evolve through the Paleozoic era, diversifying into a wide variety of forms. Many fish of the Paleozoic developed external armor that protected them from predators. The first fish with jaws appeared in the Silurian period, after which many (such as sharks) became formidable marine predators rather than just the prey of arthropods. Mos ...
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Fish Of Western Asia
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of living fish species are ray-finned fish, belonging to the class Actinopterygii, with around 99% of those being teleosts. The earliest organisms that can be classified as fish were soft-bodied chordates that first appeared during the Cambrian period. Although they lacked a true spine, they possessed notochords which allowed them to be more agile than their invertebrate counterparts. Fish would continue to evolve through the Paleozoic era, diversifying into a wide variety of forms. Many fish of the Paleozoic developed external armor that protected them from predators. The first fish with jaws appeared in the Silurian period, after which many (such as sharks) became formidable marine predators rather than just the prey of arthropods. Mos ...
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Fish Of The Caspian Sea
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of living fish species are ray-finned fish, belonging to the class Actinopterygii, with around 99% of those being teleosts. The earliest organisms that can be classified as fish were soft-bodied chordates that first appeared during the Cambrian period. Although they lacked a true spine, they possessed notochords which allowed them to be more agile than their invertebrate counterparts. Fish would continue to evolve through the Paleozoic era, diversifying into a wide variety of forms. Many fish of the Paleozoic developed external armor that protected them from predators. The first fish with jaws appeared in the Silurian period, after which many (such as sharks) became formidable marine predators rather than just the prey of arthropods. Most f ...
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Proterorhinus
''Proterorhinus'' is a genus of fishes, known as the tubenose gobies. These gobies, gobiid fish are native to Eurasia where they occur in the region of the Caspian Sea, Caspian and Black Sea, Black seas, inhabiting marine, brackish and fresh waters. The species ''Proterorhinus semilunaris'' (previously referred to as ''P. marmoratus'') was introduced to the St. Clair River in Michigan during the late 1990s. Until recently, the genus was considered monotypic, comprising only the tubenose goby (''Proterorhinus marmoratus''). Following molecular and further morphological investigations it has been split into several taxa, with distinct distributions in marine vs. fresh waters and in the Black Sea vs. Caspian Sea basins. Species There are five recognized species in this genus: * ''Proterorhinus marmoratus'' (Peter Simon Pallas, Pallas, 1814) (tubenose goby) * ''Proterorhinus nasalis'' (Filippo de Filippi, De Filippi, 1863) (eastern tubenose goby) * ''Proterorhinus semilunaris'' (Joh ...
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Proterorhinus Semipellucidus
''Proterorhinus semipellucidus '' is a species of gobiid fish, a tubenose goby originally described from the Gharasu River near Gorgan Bay of the Caspian Sea in Iran. Following the systematic decomposition of the tubenose gobies it was suggested to be a more widespread and invasive taxon distributed in the fresh waters of the Caspian Sea basin.Neilson M.E., Stepien C.A. (2009) Evolution and phylogeography of the tubenose goby genus ''Proterorhinus'' (Gobiidae: Teleostei): evidence for new cryptic species. Biological Journal of the Linnean Society, 96(3): 664–68/ref> It may be the same species as that known as '' Proterorhinus nasalis''.Brian W. CoadFreshwater Fishes of Iran(accessed 22 Feb 2015) References External links * Eschmeyer, William N. (2015semipellucidus, Gobius In ''Catalog of Fishes Catalog of Fishes is a comprehensive on-line database and reference work on the scientific names of fish species and genera. It is global in its scope and is hosted by the Cali ...
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Fish Measurement
Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish measured from the tip of the snout to the posterior end of the last vertebra or to the posterior end of the midlateral portion of the hypural plate. Simply put, this measurement excludes the length of the caudal (tail) fin. * Total length (TL) is the length of a fish measured from the tip of the snout to the tip of the longer lobe of the caudal fin, usually measured with the lobes compressed along the midline. It is a straight-line measure, not measured over the curve of the body. Standard length measurements are used with Teleostei (most bony fish), while total length measurements are used with Myxini (hagfish), Petromyzontiformes (lampreys), and (usually) Elasmobranchii (sharks and rays), as well as some other fishes. Total length meas ...
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Invasive Species
An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native species that become harmful to their native environment after human alterations to its food webfor example the purple sea urchin (''Strongylocentrotus purpuratus'') which has decimated kelp forests along the northern California coast due to overharvesting of its natural predator, the California sea otter (''Enhydra lutris''). Since the 20th century, invasive species have become a serious economic, social, and environmental threat. Invasion of long-established ecosystems by organisms is a natural phenomenon, but human-facilitated introductions have greatly increased the rate, scale, and geographic range of ...
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Maurice Kottelat
Maurice Kottelat (born 16 July 1957 in Delémont, SwitzerlandCommissioners: Dr Maurice Kottelat
International Commission on Zoological Nomenclature (accessed 2014)
) is a specializing in Eurasian freshwater fishes. Kottelat obtained a License in Sciences at the in 1987(outdated link:

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Caspian Sea
The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia, south of the fertile plains of Southern Russia in Eastern Europe, and north of the mountainous Iranian Plateau of Western Asia. It covers a surface area of (excluding the highly saline lagoon of Garabogazköl to its east) and a volume of . It has a salinity of approximately 1.2% (12 g/L), about a third of the salinity of average seawater. It is bounded by Kazakhstan to the northeast, Russia to the northwest, Azerbaijan to the southwest, Iran to the south, and Turkmenistan to the southeast. The sea stretches nearly from north to south, with an average width of . Its gross coverage is and the surface is about below sea level. Its main freshwater inflow, Europe's longest river, the Volga, enters at the shallow north end. Two deep ...
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Sea Of Azov
The Sea of Azov ( Crimean Tatar: ''Azaq deñizi''; russian: Азовское море, Azovskoye more; uk, Азовське море, Azovs'ke more) is a sea in Eastern Europe connected to the Black Sea by the narrow (about ) Strait of Kerch, and is sometimes regarded as a northern extension of the Black Sea. The sea is bounded by Russia on the east, by Ukraine on the northwest and southwest. The sea is largely affected by the inflow of the Don, Kuban, and other rivers, which bring sand, silt, and shells, which in turn form numerous bays, limans, and narrow spits. Because of these deposits, the sea bottom is relatively smooth and flat with the depth gradually increasing toward the middle. Also, due to the river inflow, water in the sea has low salinity and a high amount of biomass (such as green algae) that affects the water colour. Abundant plankton result in unusually high fish productivity. The sea shores and spits are low; they are rich in vegetation and bird colonie ...
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