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Progradungula Otwayensis
''Progradungula otwayensis'', commonly known as the odd-clawed spider, is a species of cribellate spider endemic to the Great Otway National Park of Victoria, Australia. It is one of only two species in the gradungulid genus ''Progradungula''. Odd-clawed spiders are medium-sized to large spiders, with an approximate body length of and a leg span of . They are lightly pigmented, from yellowish brown to light mauvish gray, with three chevron markings on the rear upper part of their abdomen. Like other gradungulids, their legs are tipped with three claws. These claws are particularly well-developed on the first and second leg pairs and are used for latching unto prey. They are ambush predators, building characteristic ladder-shaped snares close to the ground. They stand head-down on these webs waiting for prey which they then scoop up into these webs to trap them. Taxonomy Odd-clawed spiders were first described by the Australian arachnologist Graham A. Milledge in 1997. They ar ...
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Graham A
Graham and Graeme may refer to: People * Graham (given name), an English-language given name * Graham (surname), an English-language surname * Graeme (surname), an English-language surname * Graham (musician) (born 1979), Burmese singer * Clan Graham, a Scottish clan * Graham baronets Fictional characters * Graham Aker, in the anime ''Gundam 00'' * Project Graham, what a human would look like to survive a car crash Places Canada * Graham, Sudbury District, Ontario * Graham Island, part of the Charlotte Island group in British Columbia * Graham Island (Nunavut), Arctic island in Nunavut United States * Graham, Alabama * Graham, Arizona * Graham, Florida * Graham, Georgia * Graham, Daviess County, Indiana * Graham, Fountain County, Indiana * Graham, Kentucky * Graham, Missouri * Graham, North Carolina * Graham, Oklahoma * Graham, Texas * Graham, Washington Elsewhere * Graham Land, Antarctica * Graham Island (Mediterranean Sea), British name for a submerged volcanic island in ...
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Pedipalp
Pedipalps (commonly shortened to palps or palpi) are the second pair of appendages of chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ("jaws") and anterior to the first pair of walking legs. Overview Pedipalps are composed of six segments or articles: the coxa, the trochanter, the femur, the short patella, the tibia, and the tarsus. In spiders, the coxae frequently have extensions called maxillae or gnathobases, which function as mouth parts with or without some contribution from the coxae of the anterior legs. The limbs themselves may be simple tactile organs outwardly resembling the legs, as in spiders, or chelate weapons ( pincers) of great size, as in scorpions. The pedipalps of Solifugae are covered in setae, but have not been studied in detail. Comparative studies of pedipalpal morphology may suggest that leg-like pedipalps are primitive in arachnids. At present, the only ...
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Calamistrum
In spiders, the calamistrum is a row of specialized leg bristles used to comb out fine bands of silk. It is only found on cribellate spiders, that is, spiders that possess the spinning organ known as the cribellum. The calamistrum and cribellum are used to form the hackled bands of silk which are characteristic of the webs of these spiders. The calamistrum is found on the upper margin of the metatarsus of the hind legs. Each bristle of the calamistrum is serrated on one side and smooth on the other. The length of a spider's calamistrum is always equal to or greater than the width of the cribellum. The ratio between calamistrum length and cribellum width varies greatly, however, even among related species. This is likely due to differences in spinning behavior and as well as differences in the size and shape of the legs and abdomen. When male cribellate spiders reach sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans it might be considered syn ...
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Raptorial
The term ''raptorial'' implies much the same as ''predatory'' but most often refers to modifications of an arthropod's foreleg that make it function for the grasping of prey while it is consumed, where the gripping surfaces are formed from the opposing faces of two successive leg segments (''see illustration''). This is distinctly different from the grasping mechanism of a structure such as a scorpion's claw (a "chela") in which one of the opposing surfaces is an articulated digit, and not a leg segment. While this is most widely known in mantises, similarly modified legs can be found in some crustaceans (e.g., mantis shrimp), and various insect families, such as Mantispidae, Belostomatidae, Nepidae, and Naucoridae (all members of these groups have raptorial forelegs). There are numerous other lineages within various insect families that have raptorial forelegs, most commonly seen in the family Reduviidae, but also including several different families of flies, and a few thrips. ...
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Arthropod Leg
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, plural ''coxae''), ''trochanter'', ''femur'' (plural ''femora''), ''tibia'' (plural ''tibiae''), ''tarsus'' (plural ''tarsi''), ''ischium'' (plural ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (plural ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments ar ...
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Progradungula Otwayensis - Somatic Characters Of The Female
''Progradungula'' is a genus of Australian large-clawed spiders that was first described by Raymond Robert Forster & Michael R. Gray in 1979. it contains only two species: '' P. carraiensis'' and '' P. otwayensis''. The name is derived from Latin ''pro'' ("before"), and the genus name ''Gradungula ''Gradungula'' is a monotypic genus of South Pacific large-clawed spiders containing the single species, ''Gradungula sorenseni''. It is only found in New Zealand. __TOC__ Etymology The genus name is derived from Latin ' "step" and ' "claw", ...'', referring to the ancient ancestry of the genus. It is the first discovered web-building cribellate spider in a "primitive" araneomorph spider family and helped establish the idea that all araneomorph spiders evolved from cribellate ancestors. These spiders have an uncommon web-making technique and prey-capturing behavior. A small (approximately 25 by 6 mm), tilting, ladder-like platform of cribellate capturing silk is supported ...
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Carapace
A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron. Crustaceans In crustaceans, the carapace functions as a protective cover over the cephalothorax (i.e., the fused head and thorax, as distinct from the abdomen behind). Where it projects forward beyond the eyes, this projection is called a rostrum (anatomy), rostrum. The carapace is Calcification, calcified to varying degrees in different crustaceans. Zooplankton within the phylum Crustacea also have a carapace. These include Cladocera, ostracods, and Isopoda, isopods, but isopods only have a developed "cephalic shield" carapace covering the head. Arachnids In arachnids, the carapace is formed by the fusion of prosomal tergites into a single Plate (animal anatomy), plate which carries the e ...
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Araneomorphae
The Araneomorphae (also called the Labidognatha) are an infraorder of spiders. They are distinguishable by chelicerae (fangs) that point diagonally forward and cross in a pinching action, in contrast to the Mygalomorphae (tarantulas and their close kin), where they point straight down. Araneomorphs comprise the vast majority of living spiders. Distinguishing characteristics Most spider species are Araneomorphae, which have fangs that face towards each other, increasing the orientations they can employ during prey capture. They have fewer book lungs (when present), and the females typically live one year. The Mygalomorphae have fangs that face towards the ground, and which are parallel to the long axis of the spider's body, thus they have only one orientation they can employ during prey capture. They have four pairs of book lungs, and the females often live many years. Image:Atrax robustus.jpg, This ''Atrax robustus'' shows the orientation of Myglamorphae fangs. Image:Che ...
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Suborder (biology)
Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. For some groups of organisms, their orders may follow ...
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Austrochiloidea
The Austrochiloidea or austrochiloids are a group of araneomorph spiders, treated as a superfamily. The taxon contains two families of eight-eyed spiders: * Austrochilidae Zapfe, 1955 * Gradungulidae Forster, 1955 Phylogeny The monophyly of the Austrochiloidea has been supported in both morphological and molecular phylogenetic studies. The position of the clade relative to two much larger groups, Haplogynae and Entelegynae The Entelegynae or entelegynes are a subgroup of araneomorph spiders, the largest of the two main groups into which the araneomorphs were traditionally divided. Females have a genital plate ( epigynum) and a "flow through" fertilization system; m ..., has varied. A summary in 2005 showed the Austrochiloidea to be basal to both groups: Two studies have placed representatives of the Austrochiloidea between the two, suggesting they have more derived characters than previously supposed: References {{Taxonbar, from=Q54482 Araneomorphae Arachnid super ...
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Superfamily (biology)
In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family, order, class, phylum, kingdom, domain. While older approaches to taxonomic classification were phenomenological, forming groups on the basis of similarities in appearance, organic structure and behaviour, methods based on genetic analysis have opened the road to cladistics. A given rank subsumes under it less general categories, that is, more specific descriptions of life forms. Above it, each rank is classified within more general categories of organisms and groups of organisms related to each other through inheritance of traits or features from common ancestors. The rank of any ''species'' and the description of its ''genus'' is ''basic''; which means that to identify a particular organism, it is usually not necessary to specify ranks other than these first two. Consider a particular ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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