Procolophon Trigoniceps
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Procolophon Trigoniceps
''Procolophon'' is a genus of lizard-like procolophonid parareptiles that first appeared in the Early Triassic (Induan) of South Africa, Brazil, and Antarctica. It persisted through the Permian–Triassic extinction event, but went extinct in the beginning of the Early Middle Triassic. The type species is ''P. trigoniceps''. History of discovery The first ''Procolophon'' fossil was discovered in the 1870s in Donnybrook, an area southwest of Pietermaritzburg in present-day Kwa-Zulu Natal of South Africa. The fossil was accessioned to Harry Seeley, who described the fossil in 1878. Numerous other fossils have been recovered since from localities across the Eastern Cape and Free State provinces of South Africa. Description ''Procolophon'' reached a length up to , and is considered to have been a small herbivore or insectivore. The skull of ''Procolophon'' is distinct because of its latero-posteriorly facing paired cheek spikes, along with spiked dermal ossicles. Paleontolog ...
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Richard Owen
Sir Richard Owen (20 July 1804 – 18 December 1892) was an English biologist, comparative anatomist and paleontologist. Owen is generally considered to have been an outstanding naturalist with a remarkable gift for interpreting fossils. Owen produced a vast array of scientific work, but is probably best remembered today for coining the word '' Dinosauria'' (meaning "Terrible Reptile" or "Fearfully Great Reptile"). An outspoken critic of Charles Darwin's theory of evolution by natural selection, Owen agreed with Darwin that evolution occurred, but thought it was more complex than outlined in Darwin's ''On the Origin of Species''. Owen's approach to evolution can be considered to have anticipated the issues that have gained greater attention with the recent emergence of evolutionary developmental biology. Owen was the first president of the Microscopical Society of London in 1839 and edited many issues of its journal – then known as ''The Microscopic Journal''. Owen also c ...
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Harry Seeley
Harry Govier Seeley (18 February 1839 – 8 January 1909) was a British paleontologist. Early life Seeley was born in London on 18 February 1839, the second son of Richard Hovill Seeley, a goldsmith, and his second wife Mary Govier. When his father was declared bankrupt, Seeley was sent to live with a family of piano makers. Between the ages of eleven and fourteen, he went to a day school and then spent the next two years learning to make pianos. He also attended lectures at the Royal School of Mines by Thomas Henry Huxley, Edward Forbes, and other notable scientists. In 1855, with the support of his uncle, Seeley began to study law but shortly gave it up to pursue a career as an actuary. In the late 1850s, he studied English and mathematics at the Working Men's College and served as a secretary for the college's museum. He also worked in the library of the British Museum, where Samuel Pickworth Woodward encouraged him to study geology. In 1859, Seeley began studies at Sidney Su ...
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Bradysaurus
''Bradysaurus'' was a large, early and common pareiasaur, the fossils of which are known from the ''Tapinocephalus'' Assemblage Zone (Capitanian age) of the South African Karoo. Along with the similarly large dinocephalia, the bradysaurs constituted the herbivorous megafauna of the late Middle Permian Period. In life they were probably slow, clumsy and inoffensive animals, that had evolved a covering of armoured scutes to protect them against their predators, the gorgonopsians. Description ''Bradysaurus'' was in length and half a tonne to a tonne in weight. The skull was large (about 42 to 48 centimeters long), broad and rounded at the front. It was coarsely sculptured and knobby, with the sutures between the bones not clearly visible. The marginal teeth were high-crowned, with only a few cusps, which is a primitive characteristic. The feet were short and broad, the phalangeal count being 2,3,3,3,2 on the fore-foot and 2,3,3,4,3 on the hind. The whole body is prot ...
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Pareiasaur
Pareiasaurs (meaning "cheek lizards") are an extinct clade of large, herbivorous parareptiles. Members of the group were armoured with scutes which covered large areas of the body. They first appeared in southern Pangea during the Middle Permian, before becoming globally distributed during the Late Permian. Pareiasaurs were the largest reptiles of the Permian, reaching sizes equivalent to those of contemporary therapsids. Pareiasaurs became extinct at the end of the Permian during the Permian-Triassic extinction event. Description Pareiasaurs ranged in size from long, and may have weighed up to . They were stocky, with short tails, small heads, robust limbs, and broad feet. The cow-sized species '' Bunostegos'', which lived 260 million years ago, is the earliest known example of a tetrapod with a fully erect posture as its legs were positioned directly under its body. Pareiasaurs were protected by bony scutes called osteoderms that were set into the skin. Their heavy skulls wer ...
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Derived (phylogenetics)
In phylogenetics, a primitive (or ancestral) character, trait, or feature of a lineage or taxon is one that is inherited from the common ancestor of a clade (or clade group) and has undergone little change since. Conversely, a trait that appears ''within'' the clade group (that is, is present in any subgroup within the clade but not all) is called advanced or derived. A clade is a group of organisms that consists of a common ancestor and all its lineal descendants. A primitive trait is the original condition of that trait in the common ancestor; advanced indicates a notable change from the original condition. These terms in biology contain no judgement about the sophistication, superiority, value or adaptiveness of the named trait. "Primitive" in biology means only that the character appeared first in the common ancestor of a clade group and has been passed on largely intact to more recent members of the clade. "Advanced" means the character has evolved within a later subgroup ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Procolophonidae
Procolophonidae is an extinct family of small, lizard-like parareptiles known from the Late Permian to Late Triassic that were distributed across Pangaea, having been reported from Europe, North America, China, South Africa, South America, Antarctica and Australia. The most primitive procolophonids were likely insectiovous or omnivorous, more derived members of the clade developed bicusped molars, and were likely herbivorous feeding on high fiber vegetation or durophagous omnivores. Many members of the group are noted for spines projecting from the quadratojugal bone of the skull, which likely served a defensive purpose as well as possibly also for display. At least some taxa were likely fossorial burrowers. While diverse during the Early and Middle Triassic, they had very low diversity during the Late Triassic, and were extinct by the beginning of the Jurassic. Phylogeny Below is a cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a dia ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, these two parts are the neurocranium and the viscerocranium ( facial skeleton) that includes the mandible as its largest bone. The skull forms the anterior-most portion of the skeleton and is a product of cephalisation—housing the brain, and several sensory structures such as the eyes, ears, nose, and mouth. In humans these sensory structures are part of the facial skeleton. Functions of the skull include protection of the brain, fixing the distance between the eyes to allow stereoscopic vision, and fixing the position of the ears to enable sound localisation of the direction and distance of sounds. In some animals, such as horned ungulates (mammals with hooves), the skull also has a defensive function by providing the mount (on the front ...
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Ossicles
The ossicles (also called auditory ossicles) are three bones in either middle ear that are among the smallest bones in the human body. They serve to transmit sounds from the air to the fluid-filled labyrinth (cochlea). The absence of the auditory ossicles would constitute a moderate-to-severe hearing loss. The term "ossicle" literally means "tiny bone". Though the term may refer to any small bone throughout the body, it typically refers to the malleus, incus, and stapes (hammer, anvil, and stirrup) of the middle ear. Structure The ossicles are, in order from the eardrum to the inner ear (from superficial to deep): the malleus, incus, and stapes, terms that in Latin are translated as "the hammer, anvil, and stirrup". * The malleus ( la, "hammer") articulates with the incus through the incudomalleolar joint and is attached to the tympanic membrane (eardrum), from which vibrational sound pressure motion is passed. * The incus ( la, "anvil") is connected to both the other bones. * ...
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Insectivore
A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores were amphibians. When they evolved 400 million years ago, the first amphibians were piscivores, with numerous sharp conical teeth, much like a modern crocodile. The same tooth arrangement is however also suited for eating animals with exoskeletons, thus the ability to eat insects is an extension of piscivory. At one time, insectivorous mammals were scientifically classified in an order called Insectivora. This order is now abandoned, as not all insectivorous mammals are closely related. Most of the Insectivora taxa have been reclassified; those that have not yet been reclassified and found to be truly related to each other remain in the order Eulipotyphla. Although individually small, insects exist in enormous numbers. Insects make u ...
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Herbivore
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthparts adapted to rasping or grinding. Horses and other herbivores have wide flat teeth that are adapted to grinding grass, tree bark, and other tough plant material. A large percentage of herbivores have mutualistic gut flora that help them digest plant matter, which is more difficult to digest than animal prey. This flora is made up of cellulose-digesting protozoans or bacteria. Etymology Herbivore is the anglicized form of a modern Latin coinage, ''herbivora'', cited in Charles Lyell's 1830 ''Principles of Geology''.J.A. Simpson and E.S.C. Weiner, eds. (2000) ''The Oxford English Dictionary'', vol. 8, p. 155. Richard Owen employed the anglicized term in an 1854 work on fossil teeth and skeletons. ''Herbivora'' is derived from Latin ''herba' ...
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