Polytoma Difficile
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Polytoma Difficile
''Polytoma'' is a genus of flagellates in the family Chlamydomonadaceae. Algae are similar to the genus ''Chlamydomonas'', but lack chlorophyll and are colorless. Although they are not photosynthetic, they are grouped with the green algae because they are phylogenetically related to, and derived from, flagellate green algae. ''Polytoma'' species are found in waters rich in organic matter. Description ''Polytoma'' is a genus of colorless, free-living chlorophytes similar in form to ''Chlamydomonas'' but lacking chlorophyll. Two flagella emanate from the anterior papilla of the cell, and cells have two contractile vacuoles at the flagellar base. The cell body can be spherical, ellipsoid, ovoid, or kidney-shaped. ''Polytoma'' possesses a leukoplast in place of a chloroplast, in which many starch grains are concentrated; there are typically no pyrenoids. Since they lack photosynthetic capability, ''Polytoma'' species are entirely saprotrophic, obtaining nutrients from decaying ...
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Christian Gottfried Ehrenberg
Christian Gottfried Ehrenberg (19 April 1795 – 27 June 1876) was a German naturalist, zoologist, comparative anatomist, geologist, and microscopist. Ehrenberg was an evangelist and was considered to be of the most famous and productive scientists of his time. Early collections The son of a judge, Christian Gottfried Ehrenberg was born in Delitzsch, near Leipzig. He first studied theology at the University of Leipzig, then medicine and natural sciences in Berlin and became a friend of the famous explorer Alexander von Humboldt. In 1818, he completed his doctoral dissertation on fungi, ''Sylvae mycologicae Berolinenses.'' In 1820–1825, on a scientific expedition to the Middle East with his friend Wilhelm Hemprich, he collected thousands of specimens of plants and animals. He investigated parts of Egypt, the Libyan Desert, the Nile valley and the northern coasts of the Red Sea, where he made a special study of the corals. Subsequently, parts of Syria, Arabia and Abyss ...
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Chlamydomonas
''Chlamydomonas'' is a genus of green algae consisting of about 150 speciesSmith, G.M. 1955 ''Cryptogamic Botany Volume 1. Algae and Fungi'' McGraw-Hill Book Company Inc of unicellular flagellates, found in stagnant water and on damp soil, in freshwater, seawater, and even in snow as "snow algae". ''Chlamydomonas'' is used as a model organism for molecular biology, especially studies of flagellar motility and chloroplast dynamics, biogenesis, and genetics. One of the many striking features of ''Chlamydomonas'' is that it contains ion channels (channelrhodopsins) that are directly activated by light. Some regulatory systems of ''Chlamydomonas'' are more complex than their homologs in Gymnosperms, with evolutionarily related regulatory proteins being larger and containing additional domains. Molecular phylogeny studies indicated that the traditional genus ''Chlamydomonas'' as defined using morphological data was polyphyletic within Volvocales. Many species were subsequently ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Polyphyletic
A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of convergent evolution. The arrangement of the members of a polyphyletic group is called a polyphyly .. ource for pronunciation./ref> It is contrasted with monophyly and paraphyly. For example, the biological characteristic of warm-bloodedness evolved separately in the ancestors of mammals and the ancestors of birds; "warm-blooded animals" is therefore a polyphyletic grouping. Other examples of polyphyletic groups are algae, C4 photosynthetic plants, and edentates. Many taxonomists aim to avoid homoplasies in grouping taxa together, with a goal to identify and eliminate groups that are found to be polyphyletic. This is often the stimulus for major revisions of the classification schemes. Researchers concerned more with ecology than with systema ...
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Circumscription (taxonomy)
In biological taxonomy, circumscription is the content of a taxon, that is, the delimitation of which subordinate taxa are parts of that taxon. If we determine that species X, Y, and Z belong in Genus A, and species T, U, V, and W belong in Genus B, those are our circumscriptions of those two genera. Another systematist might determine that T, U, V, W, X, Y, and Z all belong in genus A. Agreement on circumscriptions is not governed by the Codes of Zoological or Botanical Nomenclature, and must be reached by scientific consensus. A goal of biological taxonomy is to achieve a stable circumscription for every taxon. This goal conflicts, at times, with the goal of achieving a natural classification that reflects the evolutionary history of divergence of groups of organisms. Balancing these two goals is a work in progress, and the circumscriptions of many taxa that had been regarded as stable for decades are in upheaval in the light of rapid developments in molecular phylogenetics ...
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Zoospore
A zoospore is a motile asexual spore that uses a flagellum for locomotion. Also called a swarm spore, these spores are created by some protists, bacteria, and fungi to propagate themselves. Diversity Flagella types Zoospores may possess one or more distinct types of flagella - tinsel or "decorated", and whiplash, in various combinations. *Tinsellated (straminipilous) flagella have lateral filaments known as mastigonemes perpendicular to their main axis, which allow for more surface area, and disturbance of the medium, giving them the property of a rudder, that is, used for steering. *Whiplash flagella are straight, to power the zoospore through its medium. Also, the "default" zoospore only has the propelling, whiplash flagella. Both tinsel and whiplash flagella beat in a sinusoidal wave pattern, but when both are present, the tinsel beats in the opposite direction of the whiplash, to give two axes of control of motility. Morphological types In eukaryotes, the four main types ...
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Pyrenoid
Pyrenoids are sub-cellular micro-compartments found in chloroplasts of many algae,Giordano, M., Beardall, J., & Raven, J. A. (2005). CO2 concentrating mechanisms in algae: mechanisms, environmental modulation, and evolution. Annu. Rev. Plant Biol., 56, 99-131. and in a single group of land plants, the hornworts.Villarreal, J. C., & Renner, S. S. (2012) Hornwort pyrenoids, carbon-concentrating structures, evolved and were lost at least five times during the last 100 million years. ''Proceedings of the National Academy of Sciences'',109(46), 1873-1887. Pyrenoids are associated with the operation of a carbon-concentrating mechanism (CCM). Their main function is to act as centres of carbon dioxide (CO2) fixation, by generating and maintaining a CO2 rich environment around the photosynthetic enzyme ribulose-1,5-bisphosphate carboxylase/oxygenase (RuBisCO). Pyrenoids therefore seem to have a role analogous to that of carboxysomes in cyanobacteria. Algae are restricted to aqueous env ...
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Chloroplast
A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, and stores it in the energy-storage molecules ATP and NADPH while freeing oxygen from water in the cells. The ATP and NADPH is then used to make organic molecules from carbon dioxide in a process known as the Calvin cycle. Chloroplasts carry out a number of other functions, including fatty acid synthesis, amino acid synthesis, and the immune response in plants. The number of chloroplasts per cell varies from one, in unicellular algae, up to 100 in plants like ''Arabidopsis'' and wheat. A chloroplast is characterized by its two membranes and a high concentration of chlorophyll. Other plastid types, such as the leucoplast and the chromoplast, contain little chlorophyll and do not carry out photosynthesis. Chloroplasts are highly dynamic—they circulat ...
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Leukoplast
Leucoplasts (λευκός leukós "white", πλαστός plastós "formed, molded") are a category of plastid and as such are organelles found in plant cells. They are non-pigmented, in contrast to other plastids such as the chloroplast. Lacking photosynthetic pigments, leucoplasts are not green and are located in non-photosynthetic tissues of plants, such as roots, bulbs and seeds. They may be specialized for bulk storage of starch, lipid or protein and are then known as amyloplasts, elaioplasts, or proteinoplasts (also called aleuroplasts) respectively. However, in many cell types, leucoplasts do not have a major storage function and are present to provide a wide range of essential biosynthetic functions, including the synthesis of fatty acids such as palmitic acid, many amino acids, and tetrapyrrole compounds such as heme. In general, leucoplasts are much smaller than chloroplasts and have a variable morphology, often described as amoeboid. Extensive networks of str ...
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Green Algae
The green algae (singular: green alga) are a group consisting of the Prasinodermophyta and its unnamed sister which contains the Chlorophyta and Charophyta/Streptophyta. The land plants (Embryophytes) have emerged deep in the Charophyte alga as sister of the Zygnematophyceae. Since the realization that the Embryophytes emerged within the green algae, some authors are starting to properly include them. The completed clade that includes both green algae and embryophytes is monophyletic and is referred to as the clade Viridiplantae and as the kingdom Plantae. The green algae include unicellular and colonial flagellates, most with two flagella per cell, as well as various colonial, coccoid and filamentous forms, and macroscopic, multicellular seaweeds. There are about 22,000 species of green algae. Many species live most of their lives as single cells, while other species form coenobia (colonies), long filaments, or highly differentiated macroscopic seaweeds. A few other organi ...
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Chlorophyll
Chlorophyll (also chlorophyl) is any of several related green pigments found in cyanobacteria and in the chloroplasts of algae and plants. Its name is derived from the Greek words , ("pale green") and , ("leaf"). Chlorophyll allow plants to absorb energy from light. Chlorophylls absorb light most strongly in the blue portion of the electromagnetic spectrum as well as the red portion. Conversely, it is a poor absorber of green and near-green portions of the spectrum. Hence chlorophyll-containing tissues appear green because green light, diffusively reflected by structures like cell walls, is less absorbed. Two types of chlorophyll exist in the photosystems of green plants: chlorophyll ''a'' and ''b''. History Chlorophyll was first isolated and named by Joseph Bienaimé Caventou and Pierre Joseph Pelletier in 1817. The presence of magnesium in chlorophyll was discovered in 1906, and was that element's first detection in living tissue. After initial work done by German chemi ...
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Chlamydomonadaceae
Chlamydomonadaceae is a family of algae within the order Chlamydomonadales.See the NCBIbr>webpage on Chlamydomonadaceae Data extracted from the Genera As accepted by WoRMS; * '' Agloë'' * ''Brachiomonas'' * ''Carteria'' * '' Cercidium'' * ''Chlainomonas'' * ''Chlamydomonas'' * '' Chlamydonephris'' * '' Chlorobrachis'' * '' Chloroceras'' * '' Chloromonas'' * '' Chlorotriangulum'' * '' Corbierea'' * '' Costachloris'' * '' Cylindromonas'' * '' Diplostauron'' * '' Furcilla'' * '' Gigantochloris'' * '' Gloeomonas'' * ''Heterochlamydomonas'' * '' Hirtusochloris'' * '' Hyalobrachion'' * '' Isococcus'' * '' Ixipapillifera'' * ''Lobochlamys'' * ''Lobomonas'' * '' Microglena'' * '' Oltmannsiella'' * ''Oogamochlamys'' * '' Parapolytoma'' * '' Peterfiella'' * '' Phyllariochloris'' * '' Polytoma'' * '' Pithiscus'' * '' Provasoliella'' * '' Pseudocarteria'' * '' Pseudofurcilla'' * '' Pyramichlamys'' * '' Rhysamphichloris'' * '' Sanguina'' * '' Selenoch ...
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