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Polystoma Integerrimum
''Polystoma integerrimum'' is a flatworm (platyhelminth) from the class Monogenea found in amphibians in Asia and Europe. It is an endoparasite of frogs and toads and has an unusual life cycle which synchronises with that of its host. Description ''P. integerrimum'' is a leaf-like flatworm that can grow to a maximum length of about . At the anterior (head) end are the mouth and a pair of suckers, and at the posterior end is the main device by which the parasite attaches to its host, the opisthaptor, with its three pairs of suckers, a pair of hooked anchors and marginal hooks. The flatworm's mouth is connected to a muscular pharynx, an oesophagus and a gut, but it has no anus. Ecology ''P. integerrimum'' inhabits the bladder of a frog or toad where it feeds on blood, mucus and the sloughed cells of its host. Unlike many species of flatworm, the adults are either male or female, with functional testes or ovaries. The eggs are produced throughout the year but are stored in the ho ...
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Flatworm
The flatworms, flat worms, Platyhelminthes, or platyhelminths (from the Greek πλατύ, ''platy'', meaning "flat" and ἕλμινς (root: ἑλμινθ-), ''helminth-'', meaning "worm") are a phylum of relatively simple bilaterian, unsegmented, soft-bodied invertebrates. Unlike other bilaterians, they are acoelomates (having no body cavity), and have no specialized circulatory and respiratory organ (anatomy), organs, which restricts them to having flattened shapes that allow oxygen and nutrients to pass through their bodies by diffusion. The digestive cavity has only one opening for both ingestion (intake of nutrients) and egestion (removal of undigested wastes); as a result, the food cannot be processed continuously. In traditional medicinal texts, Platyhelminthes are divided into Turbellaria, which are mostly non-parasitic animals such as planarians, and three entirely parasitic groups: Cestoda, Trematoda and Monogenea; however, since the turbellarians have since been prove ...
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Monogenea
Monogeneans are a group of ectoparasitic flatworms commonly found on the skin, gills, or fins of fish. They have a direct lifecycle and do not require an intermediate host. Adults are hermaphrodites, meaning they have both male and female reproductive structures.L.A. Tubbsa et al. (2005). "Effects of temperature on fecundity in vitro, egg hatching and reproductive development of ''Benedenia seriolae'' and ''Zeuxapta seriolae'' (Monogenea) parasitic on yellowtail kingfish Seriola lalandi". ''International Journal for Parasitology''(35), 315–327. Some monogeneans are oviparous (egg-laying) and some are viviparous (live-bearing). Oviparous varieties release eggs into the water. Viviparous varieties release larvae, which immediately attach to another host. The genus ''Gyrodactylus'' is an example of a viviparous variety, while the genus ''Dactylogyrus'' is an example of an oviparous variety. Signs and symptoms Freshwater fish that become infected with this parasite become let ...
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Amphibian
Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial animal, terrestrial, fossorial, arboreal or freshwater aquatic ecosystems. Thus amphibians typically start out as larvae living in water, but some species have developed behavioural adaptations to bypass this. The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards but, along with mammals and birds, reptiles are amniotes and do not require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in re ...
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Parasitism
Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of life. The entomologist E. O. Wilson has characterised parasites as "predators that eat prey in units of less than one". Parasites include single-celled protozoans such as the agents of malaria, sleeping sickness, and amoebic dysentery; animals such as hookworms, lice, mosquitoes, and vampire bats; fungi such as Armillaria mellea, honey fungus and the agents of ringworm; and plants such as mistletoe, dodder, and the Orobanchaceae, broomrapes. There are six major parasitic Behavioral ecology#Evolutionarily stable strategy, strategies of exploitation of animal hosts, namely parasitic castration, directly transmitted parasitism (by contact), wikt:trophic, trophicallytransmitted parasitism (by being eaten), Disease vector, vector-transmitted paras ...
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Frog
A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" ''Triadobatrachus'' is known from the Early Triassic of Madagascar, but molecular clock, molecular clock dating suggests their split from other amphibians may extend further back to the Permian, 265 Myr, million years ago. Frogs are widely distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforest. Frogs account for around 88% of extant amphibian species. They are also one of the five most diverse vertebrate orders. Warty frog species tend to be called toads, but the distinction between frogs and toads is informal, not from Taxonomy (biology), taxonomy or evolutionary history. An adult frog has a stout body, protruding eyes, anteriorly-attached tongue, limb ...
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Biological Life Cycle
In biology, a biological life cycle (or just life cycle or lifecycle when the biological context is clear) is a series of changes in form that an organism undergoes, returning to the starting state. "The concept is closely related to those of the life history, development and ontogeny, but differs from them in stressing renewal." Transitions of form may involve growth, asexual reproduction, or sexual reproduction. In some organisms, different "generations" of the species succeed each other during the life cycle. For plants and many algae, there are two multicellular stages, and the life cycle is referred to as alternation of generations. The term life history is often used, particularly for organisms such as the red algae which have three multicellular stages (or more), rather than two.Dixon, P.S. 1973. ''Biology of the Rhodophyta.'' Oliver & Boyd. Life cycles that include sexual reproduction involve alternating haploid (''n'') and diploid (2''n'') stages, i.e., a change of pl ...
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Opisthaptor
The haptor is the attachment organ of the monogeneans, a group of parasitic Platyhelminthes. The haptor is sometimes called opisthaptor (from ''opistho-'': behind) to emphasize that it is located in the posterior part of the body, and to differentiate it from the prohaptor (from ''pro-'': in front), a structure including glands located at the anterior part of the body. According to Yamaguti (1963), the chief adhesive organ of the monogeneans, the haptor, is posterior, more or less discoid, muscular, may be divided into alveoli or loculi, is usually provided with anchors, has nearly always marginal larval hooklets, or is in a reduced form with anchors. The haptor may consist of symmetrical or asymmetrical, sessile or pedunculate, muscular suckers or clamps with or without supporting sclerites; accessory adhesive organs may be present in form of armed plaques, lappets or appendices. The structure of the haptor is different in the two major groups constituting the Monogenea, nam ...
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Larvae
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The larva's appearance is generally very different from the adult form (''e.g.'' caterpillars and butterflies) including different unique structures and organs that do not occur in the adult form. Their diet may also be considerably different. Larvae are frequently adapted to different environments than adults. For example, some larvae such as tadpoles live almost exclusively in aquatic environments, but can live outside water as adult frogs. By living in a distinct environment, larvae may be given shelter from predators and reduce competition for resources with the adult population. Animals in the larval stage will consume food to fuel their transition into the adult form. In some organisms like polychaetes and barnacles, adults are immobil ...
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Tadpole
A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found in adult amphibians such as a lateral line, gills and swimming tails. As they undergo metamorphosis, they start to develop functional lungs for breathing air, and the diet of tadpoles changes drastically. A few amphibians, such as some members of the frog family Brevicipitidae, undergo direct development i.e., they do not undergo a free-living larval stage as tadpoles instead emerging from eggs as fully formed "froglet" miniatures of the adult morphology. Some other species hatch into tadpoles underneath the skin of the female adult or are kept in a pouch until after metamorphosis. Having no hard skeletons, it might be expected that tadpole fossils would not exist. However, traces of biofilms have been preserved and fossil tadpoles have ...
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Metamorphosis
Metamorphosis is a biological process by which an animal physically develops including birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some insects, fish, amphibians, mollusks, crustaceans, cnidarians, echinoderms, and tunicates undergo metamorphosis, which is often accompanied by a change of nutrition source or behavior. Animals can be divided into species that undergo complete metamorphosis (" holometaboly"), incomplete metamorphosis ("hemimetaboly"), or no metamorphosis (" ametaboly"). Scientific usage of the term is technically precise, and it is not applied to general aspects of cell growth, including rapid growth spurts. Generally organisms with a larva stage undergo metamorphosis, and during metamorphosis the organism loses larval characteristics. References to "metamorphosis" in mammals are imprecise and only colloquial, but historically idealist ideas of transformation ...
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Cloaca
In animal anatomy, a cloaca ( ), plural cloacae ( or ), is the posterior orifice that serves as the only opening for the digestive, reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles and birds, and a few mammals ( monotremes, tenrecs, golden moles, and marsupial moles), have this orifice, from which they excrete both urine and feces; this is in contrast to most placental mammals, which have two or three separate orifices for evacuation. Excretory openings with analogous purpose in some invertebrates are also sometimes referred to as cloacae. Mating through the cloaca is known as cloacal copulation, commonly referred to as cloacal kiss. The cloacal region is also often associated with a secretory organ, the cloacal gland, which has been implicated in the scent-marking behavior of some reptiles, marsupials, amphibians, and monotremes. Etymology The word is from the Latin verb ''cluo'', "(I) cleanse", thus the noun ''cloaca'', ...
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Neoteny
Neoteny (), also called juvenilization,Montagu, A. (1989). Growing Young. Bergin & Garvey: CT. is the delaying or slowing of the physiological, or somatic, development of an organism, typically an animal. Neoteny is found in modern humans compared to other primates. In progenesis or paedogenesis, sexual development is accelerated. Both neoteny and progenesis result in paedomorphism (as having the form typical of children) or paedomorphosis (changing towards forms typical of children), a type of heterochrony. It is the retention in adults of traits previously seen only in the young. Such retention is important in evolutionary biology, domestication and evolutionary developmental biology. Some authors define paedomorphism as the retention of larval traits, as seen in salamanders.Schell, S. C. ''Handbook of Trematodes of North America North of Mexico'', 1985, pg. 22 History and etymology The origins of the concept of neoteny have been traced to the Bible (as argued by Ashley Monta ...
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