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Plotopterid
Plotopteridae is an extinct family of flightless seabirds with uncertain placement, generally considered as member of order Suliformes. They exhibited remarkable convergent evolution with the penguins, particularly with the now extinct giant penguins. That they lived in the North Pacific, the other side of the world from the penguins, has led to them being described at times as the Northern Hemisphere's penguins, though they were not closely related. More recent studies have shown, however, that the shoulder-girdle, forelimb and sternum of plotopterids differ significantly from those of penguins, so comparisons in terms of function may not be entirely accurate. Plotopterids are regarded as closely related to Anhingidae (darters) and Phalacrocoracidae (cormorants). On the other hand, there is a theory that this group may have a common ancestor with penguins due to the similarity of forelimb and brain morphology. However, the endocast morphology of stem group Sphenisciformes di ...
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Phocavis
''Phocavis'' is an extinct genus of flightless seabird, belonging to the family Plotopteridae, and distantly related with modern cormorants. Its fossils, found in the Keasey Formation in Oregon, are dated from the Late Eocene. History and taxonomy The holotype of ''Phocavis'', LACM 123897, an isolated right tarsometatarsus, was collected in Late Eocene rocks belonging to the Keasey Formation, near Vernonia, Oregon, by James L. Goedert in 1979. The fossil was only described in 1988 as a new genus and species of Plotopteridae Plotopteridae is an extinct family of flightless seabirds with uncertain placement, generally considered as member of order Suliformes. They exhibited remarkable convergent evolution with the penguins, particularly with the now extinct giant peng ..., ''Phocavis maritimus''. Due to the lack of existing material on the two described genera of North American plotopterids, ''Tonsala'' and ''Plotopterum'', the validity of the genus was only assumed due to the ...
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Penguin
Penguins (order (biology), order List of Sphenisciformes by population, Sphenisciformes , family (biology), family Spheniscidae ) are a group of Water bird, aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adapted for life in the water, penguins have Countershading, countershaded dark and white plumage and flippers for swimming. Most penguins feed on krill, fish, squid and other forms of sea life which they catch with their bills and swallow it whole while swimming. A penguin has a spiny tongue and powerful jaws to grip slippery prey. They spend roughly half of their lives on land and the other half in the sea. The largest living species is the emperor penguin (''Aptenodytes forsteri''): on average, adults are about tall and weigh . The smallest penguin species is the Little penguin, little blue penguin (''Eudyptula minor''), also known as the fairy penguin, whic ...
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Penguin
Penguins (order (biology), order List of Sphenisciformes by population, Sphenisciformes , family (biology), family Spheniscidae ) are a group of Water bird, aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adapted for life in the water, penguins have Countershading, countershaded dark and white plumage and flippers for swimming. Most penguins feed on krill, fish, squid and other forms of sea life which they catch with their bills and swallow it whole while swimming. A penguin has a spiny tongue and powerful jaws to grip slippery prey. They spend roughly half of their lives on land and the other half in the sea. The largest living species is the emperor penguin (''Aptenodytes forsteri''): on average, adults are about tall and weigh . The smallest penguin species is the Little penguin, little blue penguin (''Eudyptula minor''), also known as the fairy penguin, whic ...
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Cormorant
Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the IOC adopted a consensus taxonomy of seven genera. The great cormorant (''Phalacrocorax carbo'') and the common shag (''Gulosus aristotelis'') are the only two species of the family commonly encountered in Britain and Ireland and "cormorant" and "shag" appellations have been later assigned to different species in the family somewhat haphazardly. Cormorants and shags are medium-to-large birds, with body weight in the range of and wing span of . The majority of species have dark feathers. The bill is long, thin and hooked. Their feet have webbing between all four toes. All species are fish-eaters, catching the prey by diving from the surface. They are excellent divers, and under water they propel themselves with their feet with help from their wings; some cormorant species have been ...
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Plotopterum
''Plotopterum'' is an extinct genus of flightless seabird of the family Plotopteridae, native to the North Pacific during the Late Oligocene and the Early Miocene. The only described species is ''Plotopterum joaquinensis''. History and Etymology The future holotype of ''Plotopterum'' was discovered in 1961 in the Pyramid Hill Sand member of the Jewett Sand Formation, in the San Joaquin Valley of California, by Dick Bishop and Ed Mitchell. The remains were presented by Bishop to the Natural History Museum of Los Angeles County in 1964. In 1969, Hildegarde Howard, at the time retired from her work of chief curator of science at the museum, described the remains, LACM 8927, the upper end of a left coracoid, as a new genus and species, ''Plotopterum joaquinensis'', which she ascribed to a new, monotypic family, Plotopteridae, due to its unique characteristics and adaptations towards swimming. In 1977, Hasegawa Yoshikazu, Hasegawa, Okumura and Okazaki described an almost complete bird ...
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Tonsala
''Tonsala'' is an extinct genus of Plotopteridae, a family of flightless seabird similar in biology with penguins, but more closely related to modern cormorants. The genus is known from terrains dated from the Late Oligocene of the State of Washington. History and etymology In 1979, Storrs L. Olson and Hasegawa Yoshikazu identified several fossilized specimens of Late Oligocene and Early Miocene birds found in the State of Washington and in Japan as members of the family Plotopteridae, but distinct enough from ''Plotopterum'' in their general anatomy to warrant their own genus. The Washington fossils, collected by Douglas Emlong in Late Oligocene terrains of the Pysht Formation, in the north of the Olympic Peninsula, were formally described the next year, 1980, by Olson himself, as the type of the new genus and species ''Tonsala hildegardae''. Olson ascribed to the genus the holotype USNM 256518, an incomplete specimen comprising a fragmentary humerus, fragments of a distal wi ...
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Copepteryx
''Copepteryx'' is an extinct genus of flightless bird of the family Plotopteridae, endemic to Japan during the Oligocene living from 28.4 to 23 mya, meaning it existed for approximately . Taxonomy ''Copepteryx'' was named by Olson and Hasegawa in 1996. Its type is ''Copepteryx hexeris''. It was assigned to ''Plotopteridae'' by Olson and Hasegawa in 1996. The name is derived from the Ancient Greek words ''Kope'' and ''pteryx'' meaning oar wing. The apparent reference to the 19th Century paleontologist Edward Drinker Cope is accidental. ''Copepteryx'' was a diving plotopterid bird which is similar to the ''Waimanu ''Waimanu'' is a genus of early penguin which lived during the Paleocene, soon after the Cretaceous–Paleogene extinction event, around 62–60 million years ago. It was about the size of an emperor penguin (1 metre). It is one of the most impor ...''. Description The bird was aquatic and likely ate fish and squid as well as shellfish. It is unknown if it had an ...
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Hokkaidornis
''Hokkaidornis'' is an extinct genus of penguin-like plotopterid from the Late Oligocene of Hokkaido, Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north .... References Pelecaniformes Oligocene birds of Asia Fossil taxa described in 2008 Fossils of Japan {{Pelecaniformes-stub ...
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North Pacific
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continents of Asia and Oceania in the west and the Americas in the east. At in area (as defined with a southern Antarctic border), this largest division of the World Ocean—and, in turn, the hydrosphere—covers about 46% of Earth's water surface and about 32% of its total surface area, larger than Earth's entire land area combined .Pacific Ocean
. '' Britannica Concise.'' 2008: Encyclopædia Britannica, Inc.
The centers of both the

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Darter
The darters, anhingas, or snakebirds are mainly tropical waterbirds in the family Anhingidae, which contains a single genus, ''Anhinga''. There are four living species, three of which are very common and widespread while the fourth is rarer and classified as near-threatened by the IUCN. The term ''snakebird'' is usually used without any additions to signify whichever of the completely allopatric species occurs in any one region. It refers to their long thin neck, which has a snake-like appearance when they swim with their bodies submerged, or when mated pairs twist it during their bonding displays. "Darter" is used with a geographical term when referring to particular species. It alludes to their manner of procuring food, as they impale fishes with their thin, pointed beak. The American darter (''A. anhinga'') is more commonly known as the anhinga. It is sometimes called "water turkey" in the southern United States; though the anhinga is quite unrelated to the wild turkey, they ar ...
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Crown Group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown gr ...
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Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope Carbon-13, 13C in the atmosphere was exceptionally low in comparison with the more common isotope Carbon-12, 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Popigai impact structure, Siberia and in what is now ...
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