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Plenocaris
''Plenocaris plena'' is a Cambrian arthropod with a bivalved carapace, and is known from the Burgess shale and Chengjiang. Originally described as a species of ''Yohoia'' by Walcott in 1912, it was placed into its own genus in 1974. The head has a pair of simple antennae. The body has 13 tergites, with trunk tergites 2 to 4 having pairs of elongate and uniramous appendages, with appendages absent from the other body segments. The body terminates with paired tail flukes. unlike waptiids, but similar to ''Synophalos,'' the tail flukes lack segmentation. 106 specimens of ''Plenocaris'' are known from the Greater Phyllopod bed, where they comprise 0.20% of the community. It has been suggested to be a member of Hymenocarina Hymenocarina is an order of extinct arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feede ..., which co ...
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Plenocaris
''Plenocaris plena'' is a Cambrian arthropod with a bivalved carapace, and is known from the Burgess shale and Chengjiang. Originally described as a species of ''Yohoia'' by Walcott in 1912, it was placed into its own genus in 1974. The head has a pair of simple antennae. The body has 13 tergites, with trunk tergites 2 to 4 having pairs of elongate and uniramous appendages, with appendages absent from the other body segments. The body terminates with paired tail flukes. unlike waptiids, but similar to ''Synophalos,'' the tail flukes lack segmentation. 106 specimens of ''Plenocaris'' are known from the Greater Phyllopod bed, where they comprise 0.20% of the community. It has been suggested to be a member of Hymenocarina Hymenocarina is an order of extinct arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feede ..., which co ...
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Hymenocarina
Hymenocarina is an order of extinct arthropods known from the Cambrian. They possess bivalved carapaces, typically with exposed posteriors. Members of the group are morphologically diverse and had a variety of ecologies, including as filter feeders and as predators. Recent research has generally considered them to be stem or crown group members of Mandibulata, due the presence of mandibles in at least some species. Taxonomy Hymenocarines are characterized by the combination of following characters: bivalved, convex carapace covering cephalothoracic region; cephalothorax bearing multisegmented antennules and rounded mandibles, alongside post-maxillular limbs with spiny, subdivided basis and endopods with well-developed terminal claws; absence of appendages between antennules and mandibles; median sclerite and lobate protrusions located between compound eyes; posterior tagma (abdomen) with ring-like segments and terminated by a pair of well-developed caudal rami. Based on th ...
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Burgess Shale Fossils
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's book '' Wonderful Life'' describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested. The fossil beds are in a series of shale layers, averaging and totalling about in thickness. These layers were deposited against the face of a high undersea limestone cliff. All these features were later raised up above current sea level during the cr ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Burgess Shale
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. The rock unit is a black shale and crops out at a number of localities near the town of Field in Yoho National Park Yoho National Park ( ) is a National Parks of Canada, national park of Canada. It is located within the Canadian Rockies, Rocky Mountains along the western slope of the Continental Divide of the Americas in southeastern British Columbia, bordered ... and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42 km to the south. History and significance The Burgess Shale was discovered by palaeontologist Charles Doolittle Walcott, Charles Walcott on 30 August 1909, towards the end of the season's fieldwork. He returned in 1910 with his sons, daughter, and wif ...
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Chengjiang
Chengjiang (; earlier Tchinkiang) is a city located in Yuxi, Yunnan Province, China, just north of Fuxian Lake. Administrative divisions Chengjiang City has 2 subdistricts and 4 townships. ;2 subdistricts * Fenglu () * Longjie () ;4 towns Chengjiang Fossil Site In evolutionary biology, and especially paleontology, Chengjiang is noted for soft-tissue fossil finds, of the Maotianshan Shales, dated less than 518 million years ago during the Cambrian explosion, which "are as spectacular as the Burgess Shale fauna, and significantly older." Also These fossils are considered one of the most important fossil finds of the 20th century. They contain an exquisite degree of detail, cover a diverse range of fauna, and are significant in attempts to understand the evolution of life on Earth. In 2012, the Changjiang Fossil Site became a UNESCO World Heritage Site. The fossils were first discovered by Henri Mansuy and Jaques Deprat who described them in 1912, the year after Charles Walc ...
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Yohoia
''Yohoia'' is an extinct genus of megacheiran arthropod from the Cambrian period that has been found as fossils in the Burgess Shale formation of British Columbia, Canada. It has been placed among the arachnomorpha, a group of arthropods that includes the chelicerates and possibly the trilobites. Fossils range in size from 7 to 23 mm. 711 specimens of ''Yohoia'' are known from the Greater Phyllopod bed, where they comprise 1.35% of the community. ''Yohoia'' is one of the "great appendage" arthropods. All taxa have a single pair of large pre-oral jointed limbs with branched spiny ends for grasping, impaling, or filtering food items. "Great appendage" arthropods have been seen as a polyphyletic group where the appendage has independently evolved, or as a Class Megacheira including ''Yohoia'' (with ''Leanchoilia'', ''Alalcomenaeus'', ''Oestokerkus'', ''Fortiforceps'', ''Jianfengia'', ''Yawunik'') defined as euarthropods, plus the radiodonts, defined as a sister group to arthro ...
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Charles Doolittle Walcott
Charles Doolittle Walcott (March 31, 1850February 9, 1927) was an American paleontologist, administrator of the Smithsonian Institution from 1907 to 1927, and director of the United States Geological Survey.Wonderful Life (book) by Stephen Jay Gould published in 1989, Chapter 4 He is famous for his discovery in 1909 of well-preserved fossils, including some of the oldest soft-part imprints, in the Burgess Shale of British Columbia, Canada. Early life Charles Doolittle Walcott was born on March 31, 1850 in New York Mills, New York. His grandfather, Benjamin S. Walcott, moved from Rhode Island in 1822. His father, also Charles Doolittle Walcott, died when Charles Jr. was only two. Walcott was the youngest of four children. He was interested in nature from an early age, collecting minerals and bird eggs and, eventually, fossils. He attended various schools in the Utica area but left at the age of eighteen without completing high school, the end of his formal education. His intere ...
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Antenna (biology)
Antennae ( antenna), sometimes referred to as "feelers", are paired appendages used for sensing in arthropods. Antennae are connected to the first one or two segments of the arthropod head. They vary widely in form but are always made of one or more jointed segments. While they are typically sensory organs, the exact nature of what they sense and how they sense it is not the same in all groups. Functions may variously include sensing touch, air motion, heat, vibration (sound), and especially smell or taste. Antennae are sometimes modified for other purposes, such as mating, brooding, swimming, and even anchoring the arthropod to a substrate. Larval arthropods have antennae that differ from those of the adult. Many crustaceans, for example, have free-swimming larvae that use their antennae for swimming. Antennae can also locate other group members if the insect lives in a group, like the ant. The common ancestor of all arthropods likely had one pair of uniramous (unbranched ...
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Tergum
A ''tergum'' (Latin for "the back"; plural ''terga'', associated adjective tergal) is the Anatomical terms of location#Dorsal and ventral, dorsal ('upper') portion of an arthropod segment other than the head. The Anatomical terms of location#Anterior and posterior, anterior edge is called the 'base' and posterior edge is called the 'apex' or 'margin'. A given tergum may be divided into hardened plates or sclerites commonly referred to as tergites. In a Thorax (insect anatomy), thoracic segment, for example, the tergum may be divided into an anterior notum and a posterior Scutellum (insect), scutellum. Lateral extensions of a tergite are known as paranota (Greek for "alongside the back") or ''carinae'' (Latin for "keel"), exemplified by the flat-backed millipedes of the order Polydesmida. Kinorhynchs have tergal and sternal plates too, though seemingly not homologous with those of arthropods. Tergo-tergal is a stridulation, stridulatory mechanism in which fine spines of the abdom ...
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Arthropod Leg
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, plural ''coxae''), ''trochanter'', ''femur'' (plural ''femora''), ''tibia'' (plural ''tibiae''), ''tarsus'' (plural ''tarsi''), ''ischium'' (plural ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (plural ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments ar ...
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Synophalos
''Synophalos'' is an extinct genus of bivalved arthropod. The genus contains a single species, ''Synophalos xynos'', found in the Cambrian Stage 3-aged Chengjiang biota of Yunnan Yunnan , () is a landlocked Provinces of China, province in Southwest China, the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is ..., China. The body has approximately 6 (possibly 7) abdominal segments, which terminate in a forked unsegmented tail. It is noted for having been found in unusual chain-like associations where up to 20 individuals were connected via the insertion of the tail into the head carapace of the preceding individual. Similar behaviour is not known in any other arthropod, living or extinct. Its purpose is unknown, though it may have served a reproductive, migratory, or defensive purpose, with the authors of the describing paper considering migration as its most like ...
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