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Paucituberculata
Paucituberculata is an order of South American marsupials. Although currently represented only by the seven living species of shrew opossums, this order was formerly much more diverse, with more than 60 extinct species named from the fossil record, particularly from the late Oligocene to early Miocene epochs. The earliest paucituberculatans date to the late Paleocene (Itaboraian South American land mammal age). The group went through a pronounced decline in the middle Miocene epoch, which resulted in the extinction of all families of this order except for the living shrew opossums (Caenolestidae). Extinct families of Paucituberculatans include Pichipilidae, Palaeothentidae, and Abderitidae. Classification It is one of two clades of Ameridelphia, a paraphyletic group; genetic studies have shown these animals to be a sister group to Australidelphia (i.e., Didelphimorphia branched off first). The Paucituberculata were once considered to be closely related to South American p ...
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Marsupials
Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a pouch. Marsupials include opossums, Tasmanian devils, kangaroos, koalas, wombats, wallabies, bandicoots, and the extinct thylacine. Marsupials represent the clade originating from the last common ancestor of extant metatherians, the group containing all mammals more closely related to marsupials than to placentals. They give birth to relatively undeveloped young that often reside in a pouch located on their mothers' abdomen for a certain amount of time. Close to 70% of the 334 extant species occur on the Australian continent (the mainland, Tasmania, New Guinea and nearby islands). The remaining 30% are found in the Americas—primarily in South America, thirteen in Central America, and one species, the Virginia opossum, in North America, nor ...
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Caenolestidae
The family Caenolestidae contains the seven surviving species of shrew opossum: small, shrew-like marsupials that are confined to the Andes mountains of South America. The order is thought to have diverged from the ancestral marsupial line very early. They were once included in the superorder but it is now known that Ameridelphia is paraphyletic, having given rise to Australidelphia, and thus could be considered an evolutionary grade. Genetic studies indicate that they are the second most basal order of marsupials, after the didelphimorphs. As recently as 20 million years ago, at least seven genera were in South America. Today, just three genera remain. They live in inaccessible forest and grassland regions of the High Andes. Shrews were entirely absent from South America until the Great American Interchange three million years ago, and are currently present only in the northwestern part of the continent. Traditionally, it was thought that shrew opossums lost ground to these and ...
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Shrew Opossum
The family (biology), family Caenolestidae contains the seven surviving species of shrew opossum: small, shrew-like marsupials that are confined to the Andes mountains of South America. The order is thought to have diverged from the ancestral marsupial line very early. They were once included in the superorder but it is now known that Ameridelphia is paraphyletic, having given rise to Australidelphia, and thus could be considered an evolutionary grade. Genetic studies indicate that they are the second most basal (phylogenetics), basal order of marsupials, after the didelphimorphs. As recently as 20 million years ago, at least seven genera were in South America. Today, just three genera remain. They live in inaccessible forest and grassland regions of the High Andes. Shrews were entirely absent from South America until the Great American Interchange three million years ago, and are currently present only in the northwestern part of the continent. Traditionally, it was thought that ...
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South American Land Mammal Age
The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Mya (unit), Ma during the Paleocene and continuing through to the Late Pleistocene (0.011 Ma). These periods are referred to as ages, stages, or intervals and were established using geographic place names where fossil materials where obtained.Flynn & Swisher, 1995 The basic unit of measure is the first/last boundary statement. This shows that the first appearance event of one taxon is known to predate the last appearance event of another. If two taxa are found in the same fossil quarry or at the same stratigraphic horizon, then their age-range zones overlap. Background South America was an island continent for much of the Cenozoic, or the "Age of Mammals". As a result, its mammals evolved in their own unique directions, as Australia and Madagascar still have today. Paleogeographic timeline A simplified paleogeography, paleogeographic timeline of South ...
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Shrew Opossums
The family Caenolestidae contains the seven surviving species of shrew opossum: small, shrew-like marsupials that are confined to the Andes mountains of South America. The order is thought to have diverged from the ancestral marsupial line very early. They were once included in the superorder but it is now known that Ameridelphia is paraphyletic, having given rise to Australidelphia, and thus could be considered an evolutionary grade. Genetic studies indicate that they are the second most basal order of marsupials, after the didelphimorphs. As recently as 20 million years ago, at least seven genera were in South America. Today, just three genera remain. They live in inaccessible forest and grassland regions of the High Andes. Shrews were entirely absent from South America until the Great American Interchange three million years ago, and are currently present only in the northwestern part of the continent. Traditionally, it was thought that shrew opossums lost ground to these and ...
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Ameridelphia
Ameridelphia is traditionally a superorder that includes all marsupials living in the Americas except for the Monito del monte (''Dromiciops''). It is now regarded as a paraphyletic group. Orders The orders within this group are listed below: * Order Didelphimorphia (108 species) ** Family Didelphidae: opossums * Order Paucituberculata (7 species) **Family Caenolestidae: shrew opossums Evolution and phylogenetics Modern marsupials are now understood to be an originally South American lineage that later reached Australia and diversified there in a massive adaptive radiation. Molecular data, including analysis of retrotransposon insertion sites in the nuclear DNA of a variety of marsupials, and the fossil evidence indicate that Ameridelphia might best be understood as an evolutionary grade. Since Didelphimorphia appears to be the basal marsupial group, it and Paucituberculata do not seem to be closest relatives. Meanwhile, the unranked clade Euaustralidelphia has been propose ...
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Caenolestes
The common shrew opossums (genus ''Caenolestes'') are members of the family Caenolestidae. They are found in Colombia, Ecuador, Peru, and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th .... The most recently discovered species is ''C. sangay''. References Shrew opossums Marsupials of South America Marsupial genera Neogene mammals of South America Quaternary mammals of South America Taxa named by Oldfield Thomas Prehistory of Ecuador Prehistory of Peru Prehistory of Venezuela {{marsupial-stub ...
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Itaboraian
The Itaboraian ( pt, Itaboraiense) age is a period within the Early Eocene geologic time (53.0–50.0 Mya (unit), Ma) epoch of the Paleogene, used more specifically with South American land mammal ages (SALMA). It follows the Riochican and precedes the Casamayoran age. Etymology This age is named after the Itaboraí Formation in the Itaboraí Basin in Itaboraí, a Rio de Janeiro metropolitan area, Greater Rio municipality in the state of Rio de Janeiro (state), Rio de Janeiro in Brazil not very far away from Niterói. Formations Fossils Correlations References Bibliography

;Itaboraí Formation * * * * * * ;Bogotá Formation * * * ;Cerrejón Formation * * * * * * * * * * * * * ;Chota Formation * * ;Las Flores Formation * * * ;Koluel Kaike Formation * ;Maíz Gordo Formation * ;Mogollón Formation * * ;Muñani Formation * ;Peñas Coloradas Formation * * ;Río Loro Formation * {{SALMA Itaboraian, Paleocene Sout ...
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Caenolestes Sangay
The eastern caenolestid (''Caenolestes sangay'') is a shrew opossum found on the eastern slopes of the Andes in southern Ecuador. Etymology The origin of the genus name (''Caenolestes'') drives from the Greek words ''kainos'' ("new") and ''lestes'' ("robber", "pirate"). The origin of the species name (''sangay'') derives from Sangay National Park, which is the largest Andean national park in Ecuador. The park itself is named after a volcano called Sangay, one of Ecuador's most active volcanoes that lies within the park. Sangay National Park was the site of the discovery of the eastern caenolestid in 2010. Taxonomy The eastern caenolestid is one of the five members of ''Caenolestes'', and is placed in the family Caenolestidae (shrew opossums). In the latter part of 20th century, scientists believed that ''Caenolestes'' is closely related to ''Lestoros'' (the Incan caenolestid). Over the years, it became clear that ''Lestoros'' is morphologically different from ''Caenolestes''. ...
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Australidelphia
Australidelphia is the superorder that contains roughly three-quarters of all marsupials, including all those native to Australasia and a single species — the monito del monte — from South America. All other American marsupials are members of the Ameridelphia. Analysis of retrotransposon insertion sites in the nuclear DNA of a variety of marsupials has shown that the South American monito del monte's lineage is the most basal of the superorder. The Australian australidelphians form a clade, for which the name Euaustralidelphia ("true Australidelphia") has been proposed (the branching order within this group is yet to be determined). The study also showed that the most basal of all marsupial orders are the other two South American groups ( Didelphimorphia and Paucituberculata, with the former probably branching first). This indicates that Australidelphia arose in South America along with the other major divisions of extant marsupials, and likely reached Australia via Antarctic ...
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Paleocene
The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name is a combination of the Ancient Greek ''palaiós'' meaning "old" and the Eocene Epoch (which succeeds the Paleocene), translating to "the old part of the Eocene". The epoch is bracketed by two major events in Earth's history. The K–Pg extinction event, brought on by Chicxulub impact, an asteroid impact and possibly volcanism, marked the beginning of the Paleocene and killed off 75% of living species, most famously the non-avian dinosaurs. The end of the epoch was marked by the Paleocene–Eocene Thermal Maximum (PETM), which was a major climatic event wherein about 2,500–4,500 gigatons of carbon were released into the atmosphere and ocean systems, causing a spike in global temperatures and ocean acidification. In the Pal ...
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