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Parringtonia Gracilis
''Parringtonia'' is an extinct genus of Triassic archosaur within the family Erpetosuchidae, known from the type species ''Parringtonia gracilis''. It is known from a single specimen, NHMUK R8646, found from the Anisian-age Manda Formation of Tanzania. This specimen, like most archosaur material from the Manda Formation, is fragmentary, including only a maxilla and a few postcranial bones. They show similarities with those of another archosaur called '' Erpetosuchus'', known from the Middle Triassic of Scotland and the eastern United States. The phylogenetic placement of ''Parringtonia'' and ''Erpetosuchus'' are uncertain; some studies placed them close to the group Crocodylomorpha, which includes all modern crocodylians and many extinct forms that diversified after the Triassic, but this relationship has more recently been questioned. Description NHMUK R8646 consists of a right maxilla or upper jaw bone, a left scapula or shoulder blade, part of what might be the ischium bone ...
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Anisian
In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from million years ago until million years ago. The Anisian Age succeeds the Olenekian Age (part of the Lower Triassic Epoch) and precedes the Ladinian Age. Stratigraphic definitions The stage and its name were established by Austrian geologists Wilhelm Heinrich Waagen and Carl Diener in 1895. The name comes from ''Anisus'', the Latin name of the river Enns. The original type locality is at Großreifling in the Austrian state of Styria. The base of the Anisian Stage (also the base of the Middle Triassic series) is sometimes laid at the first appearance of conodont species '' Chiosella timorensis'' in the stratigraphic record. Other stratigraphers prefer to use the base of magnetic chronozone MT1n. There is no accepted global reference profile for the base, but one ( GSSP or golden spike) was proposed at a flank of the mountain Deşli Caira in the Roman ...
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Dorsal Vertebra
In vertebrates, thoracic vertebrae compose the middle segment of the vertebral column, between the cervical vertebrae and the lumbar vertebrae. In humans, there are twelve thoracic vertebrae and they are intermediate in size between the cervical and lumbar vertebrae; they increase in size going towards the lumbar vertebrae, with the lower ones being much larger than the upper. They are distinguished by the presence of facets on the sides of the bodies for articulation with the heads of the ribs, as well as facets on the transverse processes of all, except the eleventh and twelfth, for articulation with the tubercles of the ribs. By convention, the human thoracic vertebrae are numbered T1–T12, with the first one (T1) located closest to the skull and the others going down the spine toward the lumbar region. General characteristics These are the general characteristics of the second through eighth thoracic vertebrae. The first and ninth through twelfth vertebrae contain certain ...
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Jugal
The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to the quadratojugal and maxilla, as well as other bones, which may vary by species. Anatomy The jugal bone is located on either side of the skull in the circumorbital region. It is the origin of several masticatory muscles in the skull. The jugal and lacrimal bones are the only two remaining from the ancestral circumorbital series: the prefrontal, postfrontal, postorbital, jugal, and lacrimal bones. During development, the jugal bone originates from dermal bone. In dinosaurs This bone is considered key in the determination of general traits in cases in which the entire skull has not been found intact (for instance, as with dinosaurs in paleontology). In some dinosaur genera the jugal also forms part of the lower margin of either the antorbital fenestra or the infratemporal fenestra, or both. Most commonly, this bone articu ...
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Antorbital Fenestra
An antorbital fenestra (plural: fenestrae) is an opening in the skull that is in front of the eye sockets. This skull character is largely associated with archosauriforms, first appearing during the Triassic Period. Among extant archosaurs, birds still possess antorbital fenestrae, whereas crocodylians have lost them. The loss in crocodylians is believed to be related to the structural needs of their skulls for the bite force and feeding behaviours that they employ.Preushscoft, H., Witzel, U. 2002. Biomechanical Investigations on the Skulls of Reptiles and Mammals. Senckenbergiana Lethaea 82:207–222.Rayfield, E.J., Milner, A.C., Xuan, V.B., Young, P.G. 2007. Functional Morphology of Spinosaur "Crocodile Mimic" Dinosaurs. JVP. 27(4):892–901. In some archosaur species, the opening has closed but its location is still marked by a depression, or fossa, on the surface of the skull called the antorbital fossa. The antorbital fenestra houses a paranasal sinus that is confluent with ...
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Synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Sister Taxa
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomic ...
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Antorbital Fenestra
An antorbital fenestra (plural: fenestrae) is an opening in the skull that is in front of the eye sockets. This skull character is largely associated with archosauriforms, first appearing during the Triassic Period. Among extant archosaurs, birds still possess antorbital fenestrae, whereas crocodylians have lost them. The loss in crocodylians is believed to be related to the structural needs of their skulls for the bite force and feeding behaviours that they employ.Preushscoft, H., Witzel, U. 2002. Biomechanical Investigations on the Skulls of Reptiles and Mammals. Senckenbergiana Lethaea 82:207–222.Rayfield, E.J., Milner, A.C., Xuan, V.B., Young, P.G. 2007. Functional Morphology of Spinosaur "Crocodile Mimic" Dinosaurs. JVP. 27(4):892–901. In some archosaur species, the opening has closed but its location is still marked by a depression, or fossa, on the surface of the skull called the antorbital fossa. The antorbital fenestra houses a paranasal sinus that is confluent with ...
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Otic Notch
Otic notches are invaginations in the posterior margin of the skull roof, one behind each orbit. Otic notches are one of the features lost in the evolution of amniotes from their tetrapod ancestors. The notches have been interpreted as part of an auditory structure and are often reconstructed as holding a tympanum similar to those seen in modern anurans. Analysis of the ''columella'' (the stapes in amphibians and reptiles) of labyrinthodonts however indicates that it did not function in transmitting low-energy vibrations, thus rendering these animals effectively deaf to airborne sound. The otic notch instead functioned as a spiracle, at least in the early forms.Laurin, M. (1996)Hearing in Stegocephalians from the Tree of Life Web Project The Tree of Life Web Project is an Internet project providing information about the diversity and phylogeny of life on Earth. This collaborative peer reviewed project began in 1995, and is written by biologists from around the world. The site ...
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Autapomorphies
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to the focal taxon (which may be a species, family or in general any clade). It can therefore be considered an apomorphy in relation to a single taxon. The word ''autapomorphy'', first introduced in 1950 by German entomologist Willi Hennig, is derived from the Greek words αὐτός, ''autos'' "self"; ἀπό, ''apo'' "away from"; and μορφή, ''morphḗ'' = "shape". Discussion Because autapomorphies are only present in a single taxon, they do not convey information about relationship. Therefore, autapomorphies are not useful to infer phylogenetic relationships. However, autapomorphy, like synapomorphy and plesiomorphy is a relative concept depending on the taxon in question. An autapomorphy at a given level may well be a synapomorphy at ...
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Pseudosuchia
Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Prior to 2011, the clade Pseudosuchia was often called Crurotarsi in reference to the crurotarsal ankle found in almost all members of the group, which traditionally included phytosaurs, ornithosuchids, and suchians. However, a major 2011 study of Triassic archosaur relations proposed that phytosaurs were not closely related to other traditional "crurotarsans", at least compared to "bird-line archosaurs" (Avemetatarsalians) such as pterosaurs and dinosaurs. As a result, the possession of a crurotarsal ankle was considered a plesiomorphic ("primitive") feature retained by pseudosuchians. Crurotarsi now refers to a broader group of reptiles including Pseudosuchia, Phytosauria, and Avemetatarsalia. Despite Pseudosuchia meaning "false crocodiles", th ...
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Neural Arch
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic irregular bone whose complex structure is composed primarily of bone, and secondarily of hyaline cartilage. They show variation in the proportion contributed by these two tissue types; such variations correlate on one hand with the cerebral/caudal rank (i.e., location within the backbone), and on the other with phylogenetic differences among the vertebrate taxa. The basic configuration of a vertebra varies, but the bone is its ''body'', with the central part of the body constituting the ''centrum''. The upper (closer to) and lower (further from), respectively, the cranium and its central nervous system surfaces of the vertebra body support attachment to the intervertebral discs. The posterior part of a vertebra forms a vertebral arch (in ...
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Glenoid Cavity
The glenoid fossa of the scapula or the glenoid cavity is a bone part of the shoulder. The word ''glenoid'' is pronounced or (both are common) and is from el, gléne, "socket", reflecting the shoulder joint's ball-and-socket form. It is a shallow, pyriform articular surface, which is located on the lateral angle of the scapula. It is directed laterally and forward and articulates with the head of the humerus; it is broader below than above and its vertical diameter is the longest. This cavity forms the glenohumeral joint along with the humerus. This type of joint is classified as a synovial, ball and socket joint. The humerus is held in place within the glenoid cavity by means of the long head of the biceps tendon. This tendon originates on the superior margin of the glenoid cavity and loops over the shoulder, bracing humerus against the cavity. The rotator cuff also reinforces this joint more specifically with the supraspinatus tendon to hold the head of the humerus in the ...
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