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Pagetia
''Pagetia'' Walcott, 1916. is a small genus of trilobite, assigned to the Eodiscinid family Pagetiidae and which had global distribution during the Middle Cambrian. The genus contains 55 currently recognized species, each with limited spatial and temporal ranges. Type species ''Pagetia bootes'' Walcott, 1916, from the middle Cambrian (late Wuliuan or base Drumian) Burgess Shale fossil Lagerstätte. The species occurs with ''Ptychagnostus praecurrens'' and derives originally from the "greater phyllopod bed" of Walcott quarry on "Fossil Ridge" between Wapta Mountain and Mount Field, approximately 5 kilometres north of Field, British Columbia in the Canadian Rocky Mountains. The presence of ''Pt. praecurrens'', indicates correlation with the Swedish ''praecurrens'' Biozone (''Baltoparadoxides pinus'' Biosubzone, A2 of Westergård 1946, pp. 98–100) in the upper part of the ''oelandicus'' Superzone. Taxonomy Order: Agnostida Salter, 1864. Suborder: Eodiscina Kobayashi, ...
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Eodiscidae
Eodiscidae is a family of agnostid trilobites that lived during the final Lower Cambrian (late Toyonian) and the Middle Cambrian. They are small or very small, and have a thorax of two or three segments. Eodiscidae includes nine genera (see box). Taxonomy The Eodiscids probably descended from the agnostids of family Yukoniidae. cited in Description Like other agnostids, the body of eodiscids is diminutive, the headshield (or cephalon) and tailshield (or pygidium) are of approximately same size, with 2 or 3 thoracic segments in-between, each consisting of a horizontal inner portion that abruptly passes into an inclined outer portion (fulcrate). The central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) is narrow, usually with paral ...
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Eodiscina
Eodiscina is trilobite suborder. The Eodiscina first developed near the end of the Lower Cambrian period (late Atdabanian) and became extinct at the end of the Middle Cambrian. Species are tiny to small, and have a thorax of two or three segments. Eodiscina includes six families classified under one superfamily, Eodiscoidea. Taxonomy The Eodiscina are mostly considered the more primitive suborder of the Agnostida, and the Agnostina the more advanced. Some scholars do not consider the Agnostina true trilobites, and consequently rejected the idea that they were related to the Eodiscina. Consequently, these scientists have proposed to elevate the group to ordinal level, which would thus be called Eodiscida Kobayashi, 1939. Origin The oldest known eodiscoid is ''Tsunyidiscus''. The glabella of ''Tsunyidiscus'' is extremely similar to that of ''Dipharus clarki'', and distinct from all other eodiscoids. ''D. clarki'' is thought to represent an immature stage of the redlichioid ''Bu ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The stu ...
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Bosistoa
''Bosistoa'' is a genus of four species of tree in the family Rutaceae endemic to eastern Australia. They have simple or compound leaves arranged in opposite pairs and bisexual flowers arranged in panicles, each flower with five sepals, five white petals and ten stamens. Description Plants in the genus ''Bosistoa'' are trees with simple or compound leaves arranged in opposite pairs and lack domatia. The flowers are bisexual, usually with five sepals fused at the base, sometimes almost for their full length, and five white petals. There are ten stamens that alternate in length. The fruit is a single or pair of follicles joined at the base with a woody exocarp. Each follicle contains a single smooth brown seed. Taxonomy The genus ''Bosistoa'' was first formally described in 1863 by George Bentham from an unpublished description by Ferdinand von Mueller and the description was published in ''Flora Australiensis''. The name ''Bosistoa'' honours the Melbourne pharmacist, Joseph Bosi ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Cambrian Trilobites
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian bi ...
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Agnostida Genera
Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician. Systematics The Agnostida are divided into two suborders — Agnostina and Eodiscina — which are then subdivided into a number of families. As a group, agnostids are isopygous, meaning their pygidium is similar in size and shape to their cephalon. Most agnostid species were eyeless. The systematic position of the order Agnostida within the class Trilobita remains uncertain, and there has been continuing ...
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Burgess Shale Fossils
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's book '' Wonderful Life'' describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested. The fossil beds are in a series of shale layers, averaging and totalling about in thickness. These layers were deposited against the face of a high undersea limestone cliff. All these features were later raised up above current sea level during the cr ...
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Selkirkia
''Selkirkia'' is a genus of predatory, tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale and Puncoviscana Formation. 142 specimens of ''Selkirkia'' are known from the Greater Phyllopod bed, where they comprise 0.27% of the community. In the Burgess Shale, 20% of the tapering, organic-walled tubes are preserved with the worm inside them, whereas the other 80% are empty (or sometimes occupied by one or more small agnostid trilobites). Whilst alive, the tubes were probably vertical, whereas trilobite-occupied tubes are horizontal. Morphology ''Selkirkia'' had a body divisible into a proboscis towards the anterior of a trunk enclosed by a tube. The proboscis would have been partially invertable and was armed with several spinules and spines, decreasing size distally overall. It was controlled by at least two sets of anterior retractor muscles. Immediately behind the proboscis was the trunk, smooth for the most part but lined with papillae tow ...
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Phyllopod Bed
The Phyllopod bed, designated by USNM locality number 35k, is the most famous fossil-bearing member of the Burgess Shale fossil ''Lagerstätte''. It was quarried by Charles Walcott from 1911–1917 (and later named Walcott Quarry), and was the source of 95% of the fossils he collected during this time; tens of thousands of soft-bodied fossils representing over 150 genera have been recovered from the Phyllopod bed alone. Stratigraphy and location The phyllopod bed is a 2.31 m thick layer of the 7 m thick Greater Phyllopod Bed, found in the Walcott Quarry on Fossil Ridge, between Wapta Mountain and Mount Field, at an elevation of around , around north of the railway town of Field, British Columbia, in the Canadian Rocky Mountains. It is adjacent to Mount Burgess, where Walcott first discovered the Burgess Shale formation. Walcott divided the bed into twelve units based on the rock type and fossil content. Certain fossil beds provide reference levels and can b ...
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Burgess Shale
The Burgess Shale is a fossil-bearing deposit exposed in the Canadian Rockies of British Columbia, Canada. It is famous for the exceptional preservation of the soft parts of its fossils. At old (middle Cambrian), it is one of the earliest fossil beds containing soft-part imprints. The rock unit is a black shale and crops out at a number of localities near the town of Field in Yoho National Park Yoho National Park ( ) is a National Parks of Canada, national park of Canada. It is located within the Canadian Rockies, Rocky Mountains along the western slope of the Continental Divide of the Americas in southeastern British Columbia, bordered ... and the Kicking Horse Pass. Another outcrop is in Kootenay National Park 42 km to the south. History and significance The Burgess Shale was discovered by palaeontologist Charles Doolittle Walcott, Charles Walcott on 30 August 1909, towards the end of the season's fieldwork. He returned in 1910 with his sons, daughter, and wif ...
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