Orovenator
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Orovenator
''Orovenator'' is an extinct genus of diapsid from Lower Permian (Artinskian stage) deposits of Oklahoma, United States. It is known from two partial skulls from the Richards Spur locality in Oklahoma. The holotype OMNH 74606 consists of a partial skull preserving snout and mandible, and the referred specimen, OMNH 74607, a partial skull preserving the skull roof, vertebrae and palatal elements. It was first named by Robert R. Reisz, Sean P. Modesto and Diane M. Scott in 2011 and the type species is ''Orovenator mayorum''. The generic name means "mountain", ''oro'', in Greek in reference to the Richards Spur locality, which was mountainous during the Permian period and "hunter", ''venator'', in Latin. The specific name honours Bill and Julie May. ''Orovenator'' is the oldest and most basal neodiapsid to date. A 2018 redescription by David Ford and Roger Benson found that ''Orovenator'' shared many similarities with varanopids, a group of reptile-like tetrapods tradit ...
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Orovenator
''Orovenator'' is an extinct genus of diapsid from Lower Permian (Artinskian stage) deposits of Oklahoma, United States. It is known from two partial skulls from the Richards Spur locality in Oklahoma. The holotype OMNH 74606 consists of a partial skull preserving snout and mandible, and the referred specimen, OMNH 74607, a partial skull preserving the skull roof, vertebrae and palatal elements. It was first named by Robert R. Reisz, Sean P. Modesto and Diane M. Scott in 2011 and the type species is ''Orovenator mayorum''. The generic name means "mountain", ''oro'', in Greek in reference to the Richards Spur locality, which was mountainous during the Permian period and "hunter", ''venator'', in Latin. The specific name honours Bill and Julie May. ''Orovenator'' is the oldest and most basal neodiapsid to date. A 2018 redescription by David Ford and Roger Benson found that ''Orovenator'' shared many similarities with varanopids, a group of reptile-like tetrapods tradit ...
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Neodiapsid
Neodiapsida is a clade, or major branch, of the reptilian family tree, typically defined as including all diapsids apart from some early primitive types known as the araeoscelidians. Modern reptiles and birds belong to the neodiapsid subclade Sauria. The oldest known neodiapsid is generally considered to be ''Orovenator'' from the Early Permian (Cisuralian) of North America. Basal-non saurian neodiaspids were ancestrally lizard-like, but basal Permian neodiapsids also include specialised swimming forms (''Hovasaurus'') the gliding lizard-like Weigeltisauridae, as well as the Triassic chameleon-like drepanosaurs. The position of the highly derived Mesozoic marine reptile groups Thalattosauria, Ichthyosauromorpha and Sauropterygia within Neodiapsida is uncertain, and they may lie within Sauria. Classification The clade Neodiapsida was given a phylogenetic definition by Laurin in 1991. He defined it as the branch-based clade containing all animals more closely related to "Youngin ...
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Richards Spur
Richards Spur is a Permian fossil locality located at the Dolese Brothers Limestone Quarry north of Lawton, Oklahoma. The locality preserves clay and mudstone fissure fills of a karst system eroded out of Ordovician limestone and dolomite, with the infilling dating to the Artinskian stage of the early Permian (Cisuralian), around 289 to 286 million years ago. Fossils of terrestrial animals are abundant and well-preserved, representing one of the most diverse Paleozoic tetrapod communities known. A common historical name for the site is Fort Sill, in reference to the nearby military base. Fossils were first reported at the quarry by workers in 1932, spurring a wave of collecting by local and international geologists. Early taxa of interest included the abundant reptile ''Captorhinus'' and microsaurs such as ''Cardiocephalus'' and ''Euryodus''. Later notable discoveries include ''Doleserpeton'' (one of the most lissamphibian-like Paleozoic tetrapods), the most diverse assortment of ...
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2011 In Paleontology
Protozoa New taxa Plants Ferns and fern allies Gymnosperms Gymnosperm research *An amplified whole plant reconstruction of the Ypresian Princeton chert pine '' Pinus arnoldii'', expanding the diagnosis to include ''P. similkameenensis'' (Miller, 1973 Events January * January 1 - The United Kingdom, the Republic of Ireland and Denmark enter the European Economic Community, which later becomes the European Union. * January 15 – Vietnam War: Citing progress in peace negotiations, U.S. ...) foliage and wood plus unnamed pollens cones found in attachment to the ''P. arnoldii'' ovulate cones is published by Klymiuk, Stockey, & Rothwell. Angiosperms Nematoda Lobopods Vetulicolians Molluscs Arthropods Fishes Amphibians Newly named lepospondyls Newly named temnospondyls Newly named lissamphibians Basal reptiles Newly named captorhinids Newly named basal diapsids Newly named ichthyosaurs Lepidosauromorphs Newly named saurosphargids New ...
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Diapsid
Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years ago during the late Carboniferous period. All diapsids other than the most primitive ones in the clade Araeoscelidia are sometimes placed into the clade Neodiapsida. The diapsids are extremely diverse, and include birds and all modern reptile groups, including turtles, which were historically thought to lie outside the group. Although some diapsids have lost either one hole (lizards), or both holes (snakes and turtles), or have a heavily restructured skull (modern birds), they are still classified as diapsids based on their ancestry. At least 17,084 species of diapsid animals are extant: 9,159 birds, and 7,925 snakes, lizards, tuatara, turtles, and crocodiles. Characteristics The name Diapsida means "two arches", and diapsids are tradition ...
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Varanopidae
Varanopidae is an extinct family of amniotes that resembled monitor lizards and may have filled a similar niche, hence the name. Typically, they are considered synapsids that evolved from an ''Archaeothyris''-like synapsid in the Late Carboniferous. However, some recent studies have recovered them being taxonomically closer to diapsid reptiles. A varanopid from the latest Middle Permian ''Pristerognathus'' Assemblage Zone is the youngest known varanopid and the last member of the "pelycosaur" group of synapsids. Description No known varanopids developed a sail like ''Dimetrodon''. The length of known varanopids, including the tail, varies from . Varanopids already showed some advanced characteristics of true pelycosaurs such as their deep, narrow, elongated skulls. Their jaws were long and their teeth were sharp. However, they were still primitive by mammalian standards. They had long tails, lizard-like bodies, and thin legs. The varanopids were mostly carnivorous, but as they ...
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Artinskian
In the geologic timescale, the Artinskian is an age or stage of the Permian. It is a subdivision of the Cisuralian Epoch or Series. The Artinskian likely lasted between and million years ago (Ma) according to the most recent revision of the International Commission on Stratigraphy (ICS) in 2022. It was preceded by the Sakmarian and followed by the Kungurian. Stratigraphy The Artinskian is named after the small Russian city of Arti (formerly ''Artinsk''), situated in the southern Ural mountains, about 200 km southwest of Yekaterinburg. The stage was introduced into scientific literature by Alexander Karpinsky in 1874. Base of the Artinskian The base of the Artinskian Stage is defined as the first appearance datum (FAD) of the conodont species '' Sweetognathus whitei'' and ''Mesogondolella bisselli''. In order to constrain this age, the ICS subcommission on Permian stratigraphy informally proposed a candidate GSSP in 2002, later followed by a formal proposal in 201 ...
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Specific Name (zoology)
In zoological nomenclature, the specific name (also specific epithet or species epithet) is the second part (the second name) within the scientific name of a species (a binomen). The first part of the name of a species is the name of the genus or the generic name. The rules and regulations governing the giving of a new species name are explained in the article species description. For example, the scientific name for humans is ''Homo sapiens'', which is the species name, consisting of two names: ''Homo'' is the " generic name" (the name of the genus) and ''sapiens'' is the "specific name". Historically, ''specific name'' referred to the combination of what are now called the generic and specific names. Carl Linnaeus, who formalized binomial nomenclature, made explicit distinctions between specific, generic, and trivial names. The generic name was that of the genus, the first in the binomial, the trivial name was the second name in the binomial, and the specific the proper term for ...
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Roger B
Roger is a given name, usually masculine, and a surname. The given name is derived from the Old French personal names ' and '. These names are of Germanic origin, derived from the elements ', ''χrōþi'' ("fame", "renown", "honour") and ', ' ("spear", "lance") (Hrōþigēraz). The name was introduced into England by the Normans. In Normandy, the Frankish name had been reinforced by the Old Norse cognate '. The name introduced into England replaced the Old English cognate '. ''Roger'' became a very common given name during the Middle Ages. A variant form of the given name ''Roger'' that is closer to the name's origin is ''Rodger''. Slang and other uses Roger is also a short version of the term "Jolly Roger", which refers to a black flag with a white skull and crossbones, formerly used by sea pirates since as early as 1723. From up to , Roger was slang for the word "penis". In ''Under Milk Wood'', Dylan Thomas writes "jolly, rodgered" suggesting both the sexual double entend ...
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Synapsid
Synapsids + (, 'arch') > () "having a fused arch"; synonymous with ''theropsids'' (Greek, "beast-face") are one of the two major groups of animals that evolved from basal amniotes, the other being the sauropsids, the group that includes reptiles and birds. The group includes mammals and every animal more closely related to mammals than to sauropsids. Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye orbit, leaving a bony arch beneath each; this accounts for their name. The distinctive temporal fenestra developed about 318 million years ago during the Late Carboniferous period, when synapsids and sauropsids diverged, but was subsequently merged with the orbit in early mammals. Traditionally, non-mammalian synapsids were believed to have evolved from reptiles, and therefore described as mammal-like reptiles in classical systematics, and primitive synapsids were also referred to as pelycosaurs, or pelycosaur-gr ...
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Tetrapod
Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct therapsids and all extant mammals). Tetrapods evolved from a clade of primitive semiaquatic animals known as the Tetrapodomorpha which, in turn, evolved from ancient lobe-finned fish (sarcopterygians) around 390 million years ago in the Middle Devonian period; their forms were transitional between lobe-finned fishes and true four-limbed tetrapods. Limbed vertebrates (tetrapods in the broad sense of the word) are first known from Middle Devonian trackways, and body fossils became common near the end of the Late Devonian but these were all aquatic. The first crown-tetrapods (last common ancestors of extant tetrapods capable of terrestrial locomotion) appeared by the very early Carboniferous, 350 million years ago. The specific aquatic ancestors ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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