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Ornithocheiroids
Eupterodactyloidea (meaning "true Pterodactyloidea") is an extinct group of pterodactyloid pterosaurs that existed from the latest Late Jurassic to the latest Late Cretaceous periods (Tithonian to Maastrichtian stages). Eupterodactyloids lived on all continents except Antarctica. Classification Eupterodactyloidea was named by S. Christopher Bennett in 1994 as an infraorder of the suborder Pterodactyloidea. Bennett defined it as an apomorphy-based clade. However, in 2010, Brian Andres re-defined the group as a stem-based taxon in his dissertation, and then formalized the definition in 2014 as all pterosaurs more closely related to ''Pteranodon longiceps'' than to ''Pterodactylus antiquus''. The slightly more exclusive group Ornithocheiroidea was re-defined in 2003 by Alexander Kellner. He defined it as the least inclusive clade containing ''Anhanguera blittersdorffi'', ''Pteranodon longiceps'', ''Dsungaripterus weii'', and ''Quetzalcoatlus northropi''. Ornithocheiroidea has often b ...
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Ornithocheiroidea
Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with fossil remains found all over the world except Antarctica. Ornithocheiroids were the most advanced group of pterosaurs, as the group includes the clade Azhdarchoidea, of which its members lived until the Maastrichtian stage of the Late Cretaceous, around 66 million years ago. Notable pterosaurs from this group include the pteranodontians ''Pteranodon'' and ''Nyctosaurus'', the ornithocheirid ''Ornithocheirus'', the anhanguerid ''Tropeognathus'', as well as the azhdarchids ''Hatzegopteryx'' and ''Quetzalcoatlus''. Classification The name Ornithocheiroidea was originally defined as an apomorphy-based taxon by Christopher Bennett in 1994. It was given a relationship-based definition in 2003 by Alexander Kellner, who defined it as the least in ...
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Tethydraco Regalis
''Tethydraco'' is a genus of pterodactyloid pterosaur that lived during the Late Cretaceous period (Maastrichtian stage) of what is now the area of present Morocco, about 66 million years ago. ''Tethydraco'' was originally assigned to the family Pteranodontidae. Subsequently described material suggested that it was an azhdarchid, and possibly synonymous with ''Phosphatodraco''. The type and only species is ''T. regalis''. Discovery and naming Since 2015, a group of paleontologists has been acquiring pterosaur fossils from commercial Moroccan fossil traders, who obtain these from workers in the phosphate mines on the Khouribga plateau, which is located within the Ouled Abdoun Basin. The purpose of this project is to determine pterosaur diversity in the latest Cretaceous. From this stage, no Konservat-Lagerstätten are known, sites combining a large variety of species with exceptional preservation. It is in such sites that the vast majority of pterosaur fossils and taxa have bee ...
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Volgadraco Bogolubovi
''Volgadraco'' ("Volga River dragon") is a genus of pterodactyloid pterosaur from the Upper Cretaceous of European Russia. ''Volgadraco'' was originally classified as an azhdarchid. However, recent studies have concluded that it may belong to either the family Nyctosauridae, or the family Pteranodontidae. ''Volgadraco'' is known from lower beak (holotype SGU, no. 46/104a) and postcranial fragments from the early Campanian-age Rybushka Formation of Saratov, Russia. The size of this animal, and the development of blood supply in the lower jaw are intermediate between older Santonian or Turonian azhdarchids like ''Azhdarcho'' and '' Bakonydraco'' and later Maastrichtian azhdarchids like ''Quetzalcoatlus''. ''Volgadraco'' was described in 2008 by Averianov, Arkhangelsky, and Pervushov. The type species is ''V. bogolubovi'', the specific name honouring Russian paleontologist Nikolai Nikolaevich Bogolubov. The authors consider the earlier named genus ''Bogolubovia'' to be a ''nomen ...
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Piksi Barbarulna
''Piksi'' is a potential pterosaurLongrich, N.R., Martill, D.M., and Andres, B. (2018). Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS Biology, 16(3): e2001663. genus containing the single species ''Piksi barbarulna'' (meaning "strange elbowed big bird", from Blackfoot ''piksi'', "big bird" or, specifically, "chicken" and Latin ''barbarus'' "strange, outlandish" + ''ulna'', elbow). It lived roughly 75 million years ago in what is now Montana, United States. Known from parts of a right wing – the humerus, ulna and radius (bone), radius bones – the only specimens found so far are housed in the Museum of the Rockies (collection number MOR 1113). The genus ''Piksi'' is monotypic at present. The fossils were found in 1991 by Gloria Jean Siebrecht in the Blackfeet Indian Reservation, namely at Bob's Vacation Site locality TM-088, Glacier County. Recovered from an old stratum of the upper Two Medicine Fo ...
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Stem-based Taxon
Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which can be a specimen or a taxon of lower rank, and a description in words. Phylogenetic nomenclature is currently regulated by the '' International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature ties names to clades, groups consisting of an ancestor and all its descendants. These groups can equivalently be called monophyletic. There are slightly different ways of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the named taxon. Listing all these organisms (i.e. providing a full circumscription) requires the full phylogenetic tree to be known. In practice, the ...
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Alamodactylus Byrdi
''Alamodactylus'' is an extinct genus of nyctosaurid pteranodontoid pterosaur known from the Late Cretaceous of Texas, southern United States. It contains a single species, ''Alamodactylus byrdi''. Discovery ''Alamodactylus'' is known solely from its holotype, SMU 76476, a partial left wing first described by Myers (2010).Myers, T.S., 2010, "Earliest occurrence of the Pteranodontidae (Archosauria: Pterosauria) in North America: New material from the Austin Group of Texas", ''Journal of Paleontology'' 84(6): 1071–1081 This specimen is crushed and consists of a left humerus, the distal end of the fourth wing metacarpal and the proximal end of the first wing phalanx of fourth digit. Other elements of the holotype first reported by Andres and Myers (2013) include a left proximal syncarpal, manual phalanx, and some fragments including four tapering processes that may represent skull bones, an articular surface that appears to be the mandible articulation of the right quadrate ...
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Nyctosauridae
Nyctosauridae (meaning "night lizards" or "bat lizards") is a family of specialized soaring pterosaurs of the late Cretaceous Period of North America, Africa, and possibly Europe. It was named in 1889 by Henry Alleyne Nicholson and Richard Lydekker.Nicholson, H.A. and Lydekker, R. (1889). ''A manual of palaeontology for the use of students: with a general introduction on the principles of palæontology, Volume II''. Blackwood, 1889. Nyctosaurids are characterized by their lack of all but the wing finger. In most pterosaurs, the hand has four fingers, with the fourth elongated to support the wing, and the remaining three are usually small, clawed, and used in walking or climbing. The lack of functional fingers in nyctosaurids may suggest that they spent almost all of their time in the air, rarely walking on the ground. Nyctosaurids also possessed a distinctively enlarged crest for muscle attachment on their upper arm bone, or humerus, the ''deltopectoral crest'', hatchet shaped li ...
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Pteranodon Sternbergi
''Geosternbergia'' is an extinct genus of pteranodontid pterodactyloid pterosaur from the Late Cretaceous geological period of North America. ''Geosternbergia'' was one of the largest pterosaur genera, and had a wingspan of up to . Discovery and history The first fossils of ''Geosternbergia'' were collected by American paleontologist George F. Sternberg in 1952 from the lower portion of the Niobrara Formation. The fossils of the animal looked similar to those of the species ''Pteranodon longiceps'', but the crests were set upright and in a slightly different position. In 1958, Sternberg and paleontologist Myrl V. Walker published a study about this peculiar find. In 1966, American paleontologist John Christian Harksen assigned the specimens found as a new species of ''Pteranodon'' called ''P. sternbergi'' due to its distinct upright crest that set it apart from ''P. longiceps''. Halsey W. Miller however, concluded a revision of the different species of ''Pteranodon'' in 197 ...
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Pteranodon Longiceps Mmartyniuk Wiki
''Pteranodon'' (); from Ancient Greek (''pteron'', "wing") and (''anodon'', "toothless") is a genus of pterosaur that included some of the largest known flying reptiles, with ''P. longiceps'' having a wingspan of . They lived during the late Cretaceous geological period of North America in present-day Kansas, Nebraska, Wyoming, South Dakota and Alabama. More fossil specimens of ''Pteranodon'' have been found than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved with nearly complete skulls and articulated skeletons. It was an important part of the animal community in the Western Interior Seaway. ''Pteranodon'' was not a dinosaur. By definition, all dinosaurs belong to the group Dinosauria; ''Pteranodon'' belongs to the group Pterosauria. Nonetheless, ''Pteranodon'' is the most famous pterosaur, frequently featured in dinosaur media and strongly associated with dinosaurs by the general public. While not dinosaurs, pterosaurs such as ...
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Pteranodontidae
The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head (most famously seen in ''Pteranodon'' itself). The spectacularly-crested ''Nyctosaurus'' is sometimes included in this family, though usually placed in its own family, the Nyctosauridae (Nicholson & Lydekker, 1889). Modern researchers differ in their use of the concept. S. Christopher Bennett and Alexander Kellner have concluded that ''Nyctosaurus'' was not a pteranodontid. In 1994 Bennett defined a clade Pteranodontidae, also including species of the Anhangueridae. However, this definition has not been accepted by other workers. Alexander Kellner, for example, named several additional species for specimens previously classified as ''Pteranodon'', and placed ''P. sternbergi'' in a distinct genus, ''Geosternbergia''. Kellner re-defined Ptera ...
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Pteranodontia
Pteranodontia is an extinct group of ornithocheiroid pterodactyloid pterosaurs that lived during the Late Cretaceous period (Coniacian to Maastrichtian stages) of North America and Africa. They were some of the most advanced pterosaurs, and possessed highly specialized cranial crests that may have served as sexual attraction, with males having a much larger crest. Classification Pteranodontia was originally named by Marsh in 1876. In 2003, it was given a phylogenetic definition by David Unwin as the common ancestor of ''Pteranodon'' and ''Nyctosaurus'' plus all its descendants. Though Marsh had originally named this group based on the shared absence of teeth in those species, most analyses show that all of the traditional "ornithocheiroid" pterosaurs are also members of this clade. Below is a cladogram showing the phylogenetic placement of this group from Andres and Myers (2013). In 2018, Longrich, Martill, and Andres revisited the classification, and made a different phylogen ...
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Pteranodontoidea
Pteranodontoidea (or pteranodontoids, from Greek meaning "toothless wings") is an extinct clade of ornithocheiroid pterosaurs from the Early to Late Cretaceous (early Valanginian to late Maastrichtian stages) of Asia, Africa, Europe, North America and South America. It was named by Alexander Wilhelm Armin Kellner in 1996. In 2003, Kellner defined the clade as a node-based taxon consisting of the last common ancestor of ''Anhanguera'', ''Pteranodon'' and all its descendants. The clade Ornithocheiroidea is sometimes considered to be the senior synonym of Pteranodontoidea, however it depends on its definition.Unwin, D. M., (2003): On the phylogeny and evolutionary history of pterosaurs. pp. 139-190. — ''in'' Buffetaut, E. & Mazin, J.-M., (eds.): ''Evolution and Palaeobiology of Pterosaurs''. Geological Society of London, Special Publications 217, London, 1-347 Brian Andres (2008, 2010, 2014) in his analyses, converts Ornithocheiroidea using the definition of Kellner (2003) to a ...
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