Opisthodontia (reptiles)
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Opisthodontia (reptiles)
Opisthodontia is a proposed clade of sphenodontian reptiles, uniting ''Opisthias'' from the Late Jurassic-earliest Cretaceous of Europe and North America with the Elienodontinae, a group of herbivorous sphenodontians known from the Late Triassic to Late Cretaceous. Description Teeth and diet Like other sphenodonts, opisthodonts had acrodont teeth which grew directly from the bone. They had one row of teeth on the lower jaw and two rows on the roof of the mouth. When processing food, their mandibular teeth would have slid between the outer (maxillary) teeth and inner (palatine) teeth. Some opisthodonts, such as '' Sphenotitan'', also had clusters of small teeth on the pterygoid at the center of the mouth roof. Opisthodont teeth were wide, numerous, and tightly-packed for grinding and shredding tough plant matter. Although wide shredding teeth are also known in a few other sphenodontians, such as ''Clevosaurus'' and ''Pelecymala'', the most diverse and long-lasting group of he ...
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Priosphenodon
''Priosphenodon'' is an extinct rhynchocephalian known from the early Late Cretaceous of Argentina. It is believed to have been a fairly large herbivore, having a longer snout than modern tuatara, with teeth specialised for shearing plant matter, like other members of Eilenodontinae. The teeth are densely packed with a cone-in-cone structure, and have prismatic enamel structure similar to those of mammals and the lizard ''Uromastyx,'' which was likely an adaptation to wear resistance in the absence of tooth replacement. Two species are known, ''P. avelesi'' and ''P. minimus''. The latter has also been considered a part of the separate genus ''Kaikaifilusaurus.'' Fossils of the genus have been found in the Candeleros and Cerro Barcino Formation The Cerro Barcino Formation (also known as the Gorro Frigio Formation) is a geological formation in South America whose strata span the Early Cretaceous to the earliest Late Cretaceous. The top age for the formation has been estimated ...
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Pelecymala
''Pelecymala'' is an extinct genus of sphenodontian reptile that lived in southwest England during the Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ... period. It has been recovered in recent studies as a primitive member of the group. References Prehistoric reptile genera Sphenodontia Fossil taxa described in 1986 {{Triassic-animal-stub ...
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Rhaetian
The Rhaetian is the latest age of the Triassic Period (in geochronology) or the uppermost stage of the Triassic System (in chronostratigraphy). It was preceded by the Norian and succeeded by the Hettangian (the lowermost stage or earliest age of the Jurassic). The base of the Rhaetian lacks a formal GSSP, though candidate sections include Steinbergkogel in Austria (since 2007) and Pignola-Abriola in Italy (since 2016). The end of the Rhaetian (and the base of the overlying Hettangian Stage) is more well-defined. According to the current ICS (International Commission on Stratigraphy) system, the Rhaetian ended ± 0.2 Ma (million years ago). In 2010, the base of the Rhaetian (i.e. the Norian-Rhaetian boundary) was voted to be defined based on the first appearance of '' Misikella posthernsteini'', a marine conodont. However, there is still much debate over the age of this boundary, as well as the evolution of ''M. posthernsteini''. The most comprehensive source of precise age ...
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Norian
The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic definitions The Norian was named after the Noric Alps in Austria. The stage was introduced into scientific literature by Austrian geologist Edmund Mojsisovics von Mojsvar in 1869. The Norian Stage begins at the base of the ammonite biozones of '' Klamathites macrolobatus'' and '' Stikinoceras kerri'', and at the base of the conodont biozones of '' Metapolygnathus communisti'' and '' Metapolygnathus primitius''. A global reference profile for the base (a GSSP) had in 2009 not yet been appointed. The top of the Norian (the base of the Rhaetian) is at the first appearance of ammonite species '' Cochloceras amoenum''. The base of the Rheatian is also close to the first appearance of conodont species '' Misikella spp.'' and '' Epigondolella mo ...
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Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, the fourth-largest country in the Americas, and the eighth-largest country in the world. It shares the bulk of the Southern Cone with Chile to the west, and is also bordered by Bolivia and Paraguay to the north, Brazil to the northeast, Uruguay and the South Atlantic Ocean to the east, and the Drake Passage to the south. Argentina is a federal state subdivided into twenty-three provinces, and one autonomous city, which is the federal capital and largest city of the nation, Buenos Aires. The provinces and the capital have their own constitutions, but exist under a federal system. Argentina claims sovereignty over the Falkland Islands, South Georgia and the South Sandwich Islands, and a part of Antarctica. The earliest recorded human prese ...
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Ankylosphenodon
''Ankylosphenodon'' is an extinct species of sphenodontian known from Tepexi de Rodriguez, Mexico. It is known from Early Cretaceous sedimentary deposits from the Tlayua formation. Lifestyle ''Ankylosphenodon'' is thought to have been an aquatic reptile due to its pachyostotic skeleton and other anatomical features often seen in aquatic animals. There is also evidence that its teeth were constantly growing throughout its lifetime, which may be indicative of herbivory, a rare characteristic among Lepidosauria The Lepidosauria (, from Greek meaning ''scaled lizards'') is a subclass or superorder of reptiles, containing the orders Squamata and Rhynchocephalia. Squamata includes snakes, lizards, and amphisbaenians. Squamata contains over 9,000 species .... References Sphenodontia Cretaceous reptiles of North America Prehistoric reptile genera {{Cretaceous-reptile-stub ...
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Sphenodontidae
Sphenodontidae is a family within the reptile group Rhynchocephalia, comprising taxa most closely related to the living tuatara of the genus ''Sphenodon''. Historically the taxa included within Sphenodontidae have varied greatly between analyses, and the group has lacked a formal definition. '' Cynosphenodon'' from the Early Jurassic of Mexico has consistently been recovered as a close relative of the tuatara in most analyses, with the clade containing the two often called Sphenodontinae. The herbivorous Eilenodontinae, otherwise considered part of Opisthodontia, is also sometimes considered part of this family as the sister group to Sphenodontinae. Sphenodontines first appeared during the Early Jurassic, and are characterised by a complete lower temporal bar caused by the fusion of the quadrate/quadratojugal and the jugal The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to t ...
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Tribe (biology)
In biology, a tribe is a taxonomic rank above genus, but below family and subfamily. It is sometimes subdivided into subtribes. By convention, all taxonomic ranks from genus upwards are capitalized, including both tribe and subtribe. In zoology, the standard ending for the name of a zoological tribe is "-ini". Examples include the tribes Caprini (goat-antelopes), Hominini (hominins), Bombini (bumblebees), and Thunnini (tunas). The tribe Hominini is divided into subtribes by some scientists; subtribe Hominina then comprises "humans". The standard ending for the name of a zoological subtribe is "-ina". In botany, the standard ending for the name of a botanical tribe is "-eae". Examples include the tribes Acalypheae and Hyacintheae. The tribe Hyacintheae is divided into subtribes, including the subtribe Massoniinae. The standard ending for the name of a botanical subtribe is "-inae". In bacteriology, the form of tribe names is as in botany, e.g., Pseudomonadeae, based on the ge ...
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Subfamily
In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoological names with "-inae". See also * International Code of Nomenclature for algae, fungi, and plants * International Code of Zoological Nomenclature * Rank (botany) * Rank (zoology) In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family, order, class, phylum, kingdom, domain. While ... Sources {{biology-stub ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Opisthodont Jaws
In rodents, incisor procumbency refers to the orientation of the upper incisor, defined by the position of the cutting edge of the incisor relative to the vertical plane of the incisors. Proodont incisors have the cutting edge in front of the vertical plane, orthodont teeth have it perpendicular to the plane, opisthodont incisors have it behind the plane, and hyper-opisthodont teeth have the cutting edge even behind the back of the alveolus of the incisor. Phyllotini are mostly opisthodont, but '' Auliscomys'' and '' Galenomys'' are orthodont and have sometimes even been described as proodont, and ''Eligmodontia'', '' Loxodontomys'', and some species of ''Calomys'' are hyper-opisthodont. ''Irenomys'', ''Reithrodon'', and '' Neotomys'', formerly classified as phyllotines, are also hyper-opisthodont.Steppan, 1995, p. 18 Oryzomyini are also mostly opisthodont, but ''Amphinectomys savamis'', ''Handleyomys fuscatus'', '' Melanomys caliginosus'', ''Mindomys hammondi'', ''Scolomys m ...
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