Nectridea
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Nectridea
Nectridea is the name of an extinct order of lepospondyl tetrapods from the Carboniferous and Permian periods, including animals such as ''Diplocaulus''. In appearance, they would have resembled modern newts or aquatic salamanders, although they are not close relatives of modern amphibians. They were characterized by long, flattened tails to aid in swimming, as well as numerous features of the vertebrae. Description Nectrideans are a diverse group of tetrapods, including the aquatic Urocordylidae, the presumably terrestrial Scincosauridae, and the bizarre horned members of Diplocaulidae (also known as Keraterpetonidae), which includes the "boomerang-headed" ''Diplocaulus'', one of the most famous genera of prehistoric amphibians (in the traditional sense of the word). By the time the earliest known nectrideans appeared in the Late Carboniferous fossil record, they had already diversified into these families, indicating that basal nectrideans are unknown. These different famili ...
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Lepospondyl
Lepospondyli is a diverse taxon of early tetrapods. With the exception of one late-surviving lepospondyl from the Late Permian of Morocco (''Diplocaulus minumus''), lepospondyls lived from the Early Carboniferous ( Mississippian) to the Early Permian and were geographically restricted to what is now Europe and North America. Five major groups of lepospondyls are known: Adelospondyli; Aïstopoda; Lysorophia; Microsauria; and Nectridea. Lepospondyls have a diverse range of body forms and include species with newt-like, eel- or snake-like, and lizard-like forms. Various species were aquatic, semiaquatic, or terrestrial. None were large (the biggest genus, the diplocaulid ''Diplocaulus'', reached a meter in length, but most were much smaller), and they are assumed to have lived in specialized ecological niches not taken by the more numerous temnospondyl amphibians that coexisted with them in the Paleozoic. Lepospondyli was named in 1888 by Karl Alfred von Zittel, who coined the name t ...
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Lepospondyli
Lepospondyli is a diverse taxon of early tetrapods. With the exception of one late-surviving lepospondyl from the Late Permian of Morocco (''Diplocaulus minumus''), lepospondyls lived from the Early Carboniferous ( Mississippian) to the Early Permian and were geographically restricted to what is now Europe and North America. Five major groups of lepospondyls are known: Adelospondyli; Aïstopoda; Lysorophia; Microsauria; and Nectridea. Lepospondyls have a diverse range of body forms and include species with newt-like, eel- or snake-like, and lizard-like forms. Various species were aquatic, semiaquatic, or terrestrial. None were large (the biggest genus, the diplocaulid ''Diplocaulus'', reached a meter in length, but most were much smaller), and they are assumed to have lived in specialized ecological niches not taken by the more numerous temnospondyl amphibians that coexisted with them in the Paleozoic. Lepospondyli was named in 1888 by Karl Alfred von Zittel, who coined the name t ...
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Scincosaurus
''Scincosaurus'' is an extinct genus of nectridean lepospondyl within the family Scincosauridae. History ''Scincosaurus crassus'' was first described by Bohemian paleontologist Antonín Frič in volume 1875 of "''Sitzungsberichte der königlichen Böhmischen Gesellschaft der Wissenschaften in Prague''", which at that time was the premiere scientific journal of Bohemia (the modern day Czech Republic). Frič's contribution to this volume was a list of Carboniferous animals he and his associates recently discovered at coal gas mines near the localities of Nýřany and Kounová. His list included short preliminary descriptions for many new genera and species of tetrapods, including '' Microbrachis, Branchiosaurus, Hyloplesion'' (at that time called ''Stelliosaurus''), and ''Sparodus.'' ''Scincosaurus crassus'' was among the new tetrapods from Nýřany, and its short description (erroneously) considered it a robust lacertilian ( lizard), possibly related to ''Sparodus''. A m ...
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Urocordylidae
The Urocordylidae are an extinct family (biology), family of nectridean lepospondyl amphibians. Urocordylids lived during the Late Carboniferous and Early Permian in what is now Europe and North America and are characterized by their very long, paddle-like tails. In life, they were probably newt-like and aquatic. Fossils have been found from Ireland, France, and the eastern United States. The family was named by English naturalist Richard Lydekker in 1889 and includes the well-known genera ''Urocordylus'' and ''Sauropleura'', as well as several others based on less-complete material. The family Urocordylidae is divided into two subfamilies, the Urocordylinae and the Sauropleurinae. The two groups are distinguished by the shapes of their skulls; urocordylines have short, blunt skulls, and sauropleurines have longer, pointed skulls. Description Urocordylids are distinguished by their elongated tails. Each tail vertebra has an upper crest of bone called a neural arch and a lower cr ...
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Diplocaulus
''Diplocaulus'' (meaning "double caul") is an extinct genus of lepospondyl amphibians which lived from the Late Carboniferous to the Late Permian of North America and Africa. ''Diplocaulus'' is by far the largest and best-known of the lepospondyls, characterized by a distinctive boomerang-shaped skull. Remains attributed to ''Diplocaulus'' have been found from the Late Permian of Morocco and represent the youngest-known occurrence of a lepospondyl. Description ''Diplocaulus'' had a stocky, salamander-like body, but was relatively large, reaching up to in length. Although a complete tail is unknown for the genus, a nearly complete articulated skeleton described in 1917 preserved a row of tail vertebrae near the head. This was construed as circumstantial evidence for a long, thin tail capable of reaching the head if the animal was curled up. Most studies since this discovery have argued that Anguilliform, anguiliform (eel-like) tail movement was the main force of locomotion uti ...
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Arizonerpeton
''Arizonerpeton'' is an extinct genus of nectridean lepospondyl. It contains a single species, ''Arizonerpeton wellsi''. It lived in what is now the Swisshelm Mountains of modern-day Arizona, United States. This locality belongs to the Black Prince Limestone Formation, which is dated to the middle Pennsylvanian sub-period of the Carboniferous period. Description ''Arizonerpeton'' is known exclusively from a collection of unusual vertebrae collected by Robert Wells, the namesake of its specific name. These vertebrae can be designated as belonging to the order Nectridea, a collection of long-tailed tetrapods ( amphibians in the broad sense) also including the famous "boomerang-headed" ''Diplocaulus''. Like other nectrideans, ''Arizonerpeton's'' vertebrae had a single main body (a pleurocentrum) fused to a plate-like neural spine jutting out of the top. The flared rims of the front and rear surfaces of the pleurocentum possesses small structures which would have formed tongue-an ...
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Tetrapod
Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct therapsids and all extant mammals). Tetrapods evolved from a clade of primitive semiaquatic animals known as the Tetrapodomorpha which, in turn, evolved from ancient lobe-finned fish (sarcopterygians) around 390 million years ago in the Middle Devonian period; their forms were transitional between lobe-finned fishes and true four-limbed tetrapods. Limbed vertebrates (tetrapods in the broad sense of the word) are first known from Middle Devonian trackways, and body fossils became common near the end of the Late Devonian but these were all aquatic. The first crown-tetrapods (last common ancestors of extant tetrapods capable of terrestrial locomotion) appeared by the very early Carboniferous, 350 million years ago. The specific aquatic ancestors ...
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Aistopoda
Aistopoda (Greek for " avingnot-visible feet") is an order of highly specialised snake-like stegocephalians known from the Carboniferous and Early Permian of Europe and North America, ranging from tiny forms only , to nearly in length. They first appear in the fossil record in the Mississippian period and continue through to the Early Permian. The skull is small but very specialised, with large orbits, and large fenestrae. The primitive form '' Ophiderpeton'' has a pattern of dermal bones in the skull similar in respects to the temnospondyls. But in the advanced genus ''Phlegethontia'' the skull is very light and open, reduced to a series of struts supporting the braincase against the lower jaw, just as in snakes, and it is possible that the aistopods filled the same ecological niches in the Paleozoic that snakes do today. They had an extremely elongated body, with up to 230 vertebrae. The vertebrae were holospondylous, having only a single ossification per segment. They l ...
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Lissamphibia
The Lissamphibia is a group of tetrapods that includes all modern amphibians. Lissamphibians consist of three living groups: the Salientia (frogs, toads, and their extinct relatives), the Caudata (salamanders, newts, and their extinct relatives), and the Gymnophiona (the limbless caecilians and their extinct relatives). A fourth group, the Allocaudata, was moderately successful, spanning 160 million years from the Middle Jurassic to the Early Pleistocene, but became extinct two million years ago. For several decades, this name has been used for a group that includes all living amphibians, but excludes all the main groups of Paleozoic tetrapods, such as Temnospondyli, Lepospondyli, Embolomeri, and Seymouriamorpha. Some scientists have concluded that all of the primary groups of modern amphibians—frogs, salamanders and caecilians—are closely related. Some writers have argued that the early Permian dissorophoid ''Gerobatrachus hottoni'' is a lissamphibian. If it is not, the earl ...
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Microsauria
Microsauria ("small lizards") is an extinct, possibly polyphyletic order of tetrapods from the late Carboniferous and early Permian periods. It is the most diverse and species-rich group of lepospondyls. Recently, Microsauria has been considered paraphyletic, as several other non-microsaur lepospondyl groups such as Lysorophia seem to be nested in it. Microsauria is now commonly used as a collective term for the grade of lepospondyls that were originally classified as members of Microsauria. The microsaurs all had short tails and small legs, but were otherwise quite varied in form. The group included lizard-like animals that were relatively well-adapted to living on dry land, burrowing forms, and others that, like the modern axolotl, retained their gills into adult life, and so presumably never left the water. Distribution Microsaur remains have been found from Europe and North America in Late Carboniferous and Early Permian localities. Most North American microsaurs have bee ...
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Diplocaulidae
The Diplocaulidae ("double cauls") is an extinct family of lepospondyl amphibians that arose during the Late Carboniferous and died out in the Late Permian. They are distinguished from other amphibians, extinct and extant, by the presence of strange, horn-like protrusions jutting out from the rear of their skulls; in some genera said protrusions gave their heads an almost boomerang-like outline. Phylogeny Below is a cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... modified from Germain (2010): References Diplocaulids Pennsylvanian first appearances Lopingian extinctions {{Lepospondyli-stub es:Keraterpetontidae ...
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Tuditanomorpha
Tuditanomorpha is a suborder of microsaur lepospondyls. Tuditanomorphs lived from the Late Carboniferous to the Early Permian and are known from North America and Europe. Tuditanomorphs have a similar pattern of bones in the skull roof. Tuditanomorphs display considerable variability, especially in body size, proportions, dentition, and presacral vertebral count. Currently there are seven families of tuditanomorphs, with two being monotypic. Tuditanids, gymnarthrids, and pantylids first appear in the Lower Pennsylvanian. Goniorhynchidae, Hapsidopareiontidae, Ostodolepidae, and Trihecatontidae appear in the Late Pennsylvanian and Early Permian. Classification Suborder Tuditanomorpha *Family Goniorhynchidae **''Rhynchonkos'' *Family Gymnarthridae **''Cardiocephalus'' **''Elfridia'' **''Euryodus'' **''Leiocephalikon'' **''Pariotichus'' **''Sparodus'' *Family Hapsidopareiontidae **''Hapsidopareion'' **''Llistrofus'' **''Ricnodon'' **''Saxonerpeton'' *Family Ostodolepidae **' ...
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