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Nochnitsa Geminidens Holotype Right Side
''Nochnitsa'' (rus.: ночница, literally – nightling) is a genus of gorgonopsian therapsid from the Kotelnich red beds of Permian Russia. It contains one species, ''Nochnitsa geminidens''. It the most basal known gorgonopsian and among the smallest members of the clade known to date. Discovery and naming ''Nochnitsa'' is known from only one specimen, the holotype KPM 310. It was found in Vanyushonki Member of the Kotelnich red beds, along the Vyatka River in Kirov Oblast, which dates to the latest Guadalupian or early Lopingian epochs. ''Nochnitsa'' is named after the Nocnitsa, a nocturnal hag-like creature from Slavic mythology. Its name was intended as a parallel to the Gorgons, similarly hag-like creatures from Greek mythology, which are the namesake of many genera within Gorgonopsia and the clade as a whole. The name also reflects the nocturnal habits inferred for the genus. The type species name, ''N. geminidens'', means "twin tooth" and refers to one of the autapo ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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Teeth
A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tearing food, for defensive purposes, to intimidate other animals often including their own, or to carry prey or their young. The roots of teeth are covered by gums. Teeth are not made of bone, but rather of multiple tissues of varying density and hardness that originate from the embryonic germ layer, the ectoderm. The general structure of teeth is similar across the vertebrates, although there is considerable variation in their form and position. The teeth of mammals have deep roots, and this pattern is also found in some fish, and in crocodilians. In most teleost fish, however, the teeth are attached to the outer surface of the bone, while in lizards they are attached to the inner surface of the jaw by one side. In cartilaginous fish, s ...
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Smilesaurus
''Smilesaurus'' is an extinct genus of gorgonopsian known from Africa. It lived during the Late Permian. It contains the single species ''S. ferox''. Description ''Smilesaurus'' was a large gorgonopsian, with a skull length of up to 31 centimeters. It is characterized by extremely long canine teeth, and has the proportionally longest canines of any gorgonopsian. Unlike other gorgonopsians, which probably hunted similarly to predatory reptiles, ''Smilesaurus'' probably was a true saber-toothed predator which hunted using similar tactics to saber-toothed cats. It can be distinguished by other rubidgeines by its lack of cranial pachyostosis and rugosoties, and by its relatively small orbits. Classification The classification of ''Smilesaurus'' has been disputed. It has often been included in Rubidgeinae, but it differs from other members of the clade considerably. Instead, it may be more closely related to ''Arctops'', a position supported by a phylogenetic analysis in 2018. Belo ...
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Lycaenops
''Lycaenops'' ("wolf-face") is a genus of carnivorous therapsids. It lived during the Middle Permian to the early Late Permian, about 260 mya, in what is now South Africa. Description ''Lycaenops'' measured about and weighed up to . Like the modern-day wolves from which it takes its name, ''Lycaenops'' had a long and slender skull, with a set of dog-like fangs set into both its upper and lower jaws. These pointed canine teeth were ideal for the use of stabbing and/or tearing at the flesh of any large prey that it came upon. ''Lycaenops'' most likely hunted small vertebrates such as reptiles and dicynodonts. ''Lycaenops'' walked and ran with its long legs held close to its body. This is a feature found in mammals, but not in more primitive amniotes, early reptiles, and synapsids such as pelycosaurs, whose legs are positioned to the sides of their bodies. The ability to move like a mammal would have given ''Lycaenops'' an advantage over other land vertebrates, since it would hav ...
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Arctognathus
''Arctognathus'' is an extinct genus of gorgonopsids that throve during the Late Permian in the Karoo basin of what is now South Africa. Discovery A carnivore, like all gorgonopsid, ''Arctognathus'' was given its name ("Bear jaw") in reference to its short and rounded snout. There is only one recognized species, ''A. curvimola''. Description It was a small gorgonopsid with a total length estimated at 1.1 m and an 18 cm skull. Classification Below is a cladogram from the phylogenetic analysis of Gebauer (2007): See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera tha ... References Bibliography * * Gorgonopsia Prehistoric therapsid genera Lopingian synapsids of Africa Fossil taxa described in 1911 Taxa named by Robert Broom Loping ...
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Gorgonops
''Gorgonops'' (from el, Γοργών 'Gorgon' and 'eye, face', literally 'Gorgon eye' or 'Gorgon face') is an extinct genus of gorgonopsian therapsids, of which it is the type genus, having lived during the Late Permian (Wuchiapingian), about 260–254 million years ago in what is now South Africa. Despite its popularity, ''Gorgonops'' is a medium-sized gorgonopsian (about long maximum), regularly confused by the general public with the more massive ''Inostrancevia'', known from Russia, due to their similar appearance and the various media that tend to refer them by the name of the group they belong rather than by their genus names, which does not help in differentiation. History of discovery The holotype of the type species, ''Gorgonops torvus'', was in 1876 one of the first therapsids described, by Richard Owen, who also coined the name "Dinosauria" on the basis of the first known dinosaur fossils. It was also used as the type for which Richard Lydekker described the family ...
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Eriphostoma
''Eriphostoma'' is an extinct genus of gorgonopsian therapsids known from the Middle Permian (middle Capitanian stage) of ''Tapinocephalus'' Assemblage Zone, South Africa. It has one known species, ''Eriphostoma microdon'', and was first named by Robert Broom in 1911. It is the oldest known gorgonopsian and among the smallest and most basal members of the clade. Description Gorgonopsians were a morphologically conservative group, and like all gorgonopsians, ''Eriphostoma'' would have been a quadrupedal predator. It was among the smaller members of the group, with a skull less than long. It had a relatively short, deep snout and large orbits compared to other gorgonopsians. Like all gorgonopsians, it had five incisors and a canine tooth on each side of the upper jaw, but it had only three small postcanine teeth in its maxilla. The palate of ''Eriphostoma'' has delta-shaped palatine bosses covered in numerous teeth, like ''Gorgonops'' and many basal therapsids, but not more der ...
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Inostrancevia
''Inostrancevia'' is an extinct genus of carnivorous therapsids, containing the largest members of gorgonopsians, predators characterized by long, saber-tooth-like canines. The various species inhabited European Russia during the Upper Tatarian (Vyatskian), a Russian regional stage equivalent to the Wuchiapingian and Changhsingian stage of the Late Permian period, dating from approximately 259 to 252.3 million years ago. The genus name was described posthumously, after the Bolshevik Revolution, by the Russian paleontologist Vladimir P. Amalitsky in 1922, in honor of geologist Aleksandr Inostrantsev. The first fossils attributed to ''I. alexandri'' are found in Arkhangelsk Oblast, near the Northern Dvina at the end of the 19th century, making it the first gorgonopsian known from Russia, the only place outside Africa where they are officially recognized. Some fossils of the species in question are among the most complete remains of gorgonopsians ever identified to date, the ...
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Pravoslavlevia
''Pravoslavlevia'' is an extinct genus of gorgonopsian therapsids that lived in the late Permian and is part of the Sokolki subcomplex of Russia. It had a skull long. The total length of the animal was about . Only one species (''P. parva'') is known. Classification The following cladogram showing the position of ''Pravoslavlevia'' within Gorgonopsia follows Kammerer and Masyutin, 2018: See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera tha ... Sources paleodb.org Gorgonopsia Prehistoric therapsid genera Extinct animals of Russia Fossil taxa described in 1953 {{paleo-therapsid-stub ...
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Sauroctonus
''Sauroctonus'' (from el, σαῦρος , 'lizard' and el, κτόνος , 'murderer') is an extinct genus of therapsids. ''Sauroctonus progressus'' was a large (2 m long) gorgonopsid that lived in the Late Permian epoch before the Permian-Triassic extinction event that wiped out many life forms on Earth (259-254 million years ago). Its fossils have been found in the Usili Formation ( Songea Group) of Tanzania and the Sokolki Assemblage Zone of the Volga Basin of Russia. Description ''Sauroctonus's'' flattened, triangular skull was about long, with a parietal eye, a primitive character, on the crown. The upper and lower jaw each contained one pair of massive canine teeth (the upper pair was larger); the other teeth were smaller, but were also sharp and pointed. In addition, minute, blunt teeth were present on the palatine bones. The lower jaw was widened to form a kind of chin. The long, lightly built, five-toed limbs bore a resemblance to mammals' limbs, but despite its ...
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Suchogorgon
''Suchogorgon'' is an extinct genus of gorgonopsid Gorgonopsia (from the Greek Gorgon, a mythological beast, and 'aspect') is an extinct clade of sabre-toothed therapsids from the Middle to Upper Permian roughly 265 to 252 million years ago. They are characterised by a long and narrow skull ...s from Russia. It was a medium-sized animal with a low, narrow skull. Its skull is dotted with small pits, which might have housed sensory organs in life. Its canines were large and flat as in most gorgonopsids. References Gorgonopsia Prehistoric therapsid genera Fossils of Russia {{paleo-therapsid-stub ...
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Viatkogorgon
''Viatkogorgon'' is a genus of gorgonopsian (a type of therapsid, the group that includes modern mammals) that lived during the Permian period in what is now Russia. The first fossil was found at the Kotelnich locality near the Vyatka River and was made the holotype of the new genus and species ''V. ivachnenkoi'' in 1999. The generic name refers to the river and the related genus ''Gorgonops''—the gorgons of Greek mythology are often referenced in the names of the group. The specific name honors the paleontologist Mikhail F. Ivakhnenko. The holotype skeleton is one of the most complete gorgonopsian specimens known and includes rarely preserved elements such as gastralia (abdominal ribs) and a sclerotic ring (a bony ring inside the eye). A larger, but poorly preserved specimen has also been assigned to the species. The holotype specimen is about long, including the long skull, making ''Viatkogorgon'' a relatively small gorgonopsian. The assigned specimen is larger, with a ...
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