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Neobalaenidae
Neobalaenidae is a family of baleen whales (suborder Mysticeti) including the extant pygmy right whale. Although traditionally considered related to balaenids, a recent phylogenetic study by Fordyce and Marx (2013) recovered the living pygmy right whale as a member of Cetotheriidae, making it the only extant cetotheriid, but not all authors agree with this argument. Taxonomy The family Neobalaenidae was long restricted to the pygmy right whale from the Southern Hemisphere due to the unusual skeletal form of the species relative to other extant mysticetes. Until the early 2010s Neobalaenidae was unknown from the fossil record despite a study by Sasaki et al. (2005) placing the divergence date of Neobalaenidae from other living baleen whales at 23 mya. Fordyce and Marx found that the pygmy right whale formed a well-supported clade with Eschrichtiidae and Balaenopteridae based on molecular data, and that, within 'cetotheres', it was most closely related to the herpetocetines ('' H ...
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Pygmy Right Whale
The pygmy right whale (''Caperea marginata'') is a species of baleen whale. It may be a member of the cetotheres, a family of baleen whales which until 2012 were thought to be extinct; ''C. marginata'' has otherwise been considered the monotypic taxon sole member of the family Neobalaenidae and is the only member of the genus ''Caperea''. First described by John Edward Gray in 1846, it is the smallest of the baleen whales, ranging between in length and in mass. Despite its name, the pygmy right whale may have more in common with the gray whale and rorquals than the bowhead and right whales. The pygmy right whale is found in temperate waters of the Southern Hemisphere, and feeds on copepods and euphausiids. Little is known about its population or social habits. Unlike most other baleen whales, it has rarely been subject to exploitation. Taxonomy During the 1839-43 voyage of James Clark Ross, naturalists found bones and baleen plates resembling a smaller version of the rig ...
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Pygmy Right Whale
The pygmy right whale (''Caperea marginata'') is a species of baleen whale. It may be a member of the cetotheres, a family of baleen whales which until 2012 were thought to be extinct; ''C. marginata'' has otherwise been considered the monotypic taxon sole member of the family Neobalaenidae and is the only member of the genus ''Caperea''. First described by John Edward Gray in 1846, it is the smallest of the baleen whales, ranging between in length and in mass. Despite its name, the pygmy right whale may have more in common with the gray whale and rorquals than the bowhead and right whales. The pygmy right whale is found in temperate waters of the Southern Hemisphere, and feeds on copepods and euphausiids. Little is known about its population or social habits. Unlike most other baleen whales, it has rarely been subject to exploitation. Taxonomy During the 1839-43 voyage of James Clark Ross, naturalists found bones and baleen plates resembling a smaller version of the rig ...
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Baleen Whale
Baleen whales (systematic name Mysticeti), also known as whalebone whales, are a parvorder of carnivorous marine mammals of the infraorder Cetacea (whales, dolphins and porpoises) which use keratinaceous baleen plates (or "whalebone") in their mouths to sieve planktonic creatures from the water. Mysticeti comprises the families Balaenidae (right and bowhead whales), Balaenopteridae (rorquals and the gray whale), and Cetotheriidae (the pygmy right whale). There are currently 16 species of baleen whales. While cetaceans were historically thought to have descended from mesonychids, molecular evidence instead supports them as a clade of even-toed ungulates (Artiodactyla). Baleen whales split from toothed whales (Odontoceti) around 34 million years ago. Baleen whales range in size from the and pygmy right whale to the and blue whale, the largest known animal to have ever existed. They are sexually dimorphic. Baleen whales can have streamlined or large bodies, depending on th ...
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Mysticeti
Baleen whales (systematic name Mysticeti), also known as whalebone whales, are a parvorder of carnivorous marine mammals of the infraorder Cetacea (whales, dolphins and porpoises) which use keratinaceous baleen plates (or "whalebone") in their mouths to sieve planktonic creatures from the water. Mysticeti comprises the families Balaenidae (right and bowhead whales), Balaenopteridae (rorquals and the gray whale), and Cetotheriidae (the pygmy right whale). There are currently 16 species of baleen whales. While cetaceans were historically thought to have descended from mesonychids, molecular evidence instead supports them as a clade of even-toed ungulates (Artiodactyla). Baleen whales split from toothed whales (Odontoceti) around 34 million years ago. Baleen whales range in size from the and pygmy right whale to the and blue whale, the largest known animal to have ever existed. They are sexually dimorphic. Baleen whales can have streamlined or large bodies, depending on th ...
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Baleen Whales
Baleen whales (systematic name Mysticeti), also known as whalebone whales, are a parvorder of carnivorous marine mammals of the infraorder Cetacea (whales, dolphins and porpoises) which use keratinaceous baleen plates (or "whalebone") in their mouths to sieve planktonic creatures from the water. Mysticeti comprises the families Balaenidae (right and bowhead whales), Balaenopteridae (rorquals and the gray whale), and Cetotheriidae (the pygmy right whale). There are currently 16 species of baleen whales. While cetaceans were historically thought to have descended from mesonychids, molecular evidence instead supports them as a clade of even-toed ungulates (Artiodactyla). Baleen whales split from toothed whales (Odontoceti) around 34 million years ago. Baleen whales range in size from the and pygmy right whale to the and blue whale, the largest known animal to have ever existed. They are sexually dimorphic. Baleen whales can have streamlined or large bodies, depending on ...
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Balaenidae
Balaenidae () is a family of whales of the parvorder Mysticeti that contains two living genera: the right whales (genus ''Eubalaena''), and in a separate genus, the closely related bowhead whale (genus ''Balaena''). Evolutionary history Baleen whales belong to a monophyletic lineage of Mysticeti. Mysticeti are large filter-feeding cetaceans that also include some of the largest animals on earth as well as some of the most critically endangered.''Kaliszewska, Z. A.J. SegerV. Rowntreeet al2005Population histories'' of right whales (Cetacea: Eubalaena) inferred from mitochondrial sequence diversities and divergences of their whale lice (Amphipoda: Cyamus)Molecular Ecology1434393456 Based on morphology and molecular data, four extant family-level clades are recognized within Mysticeti: Balaenidae (bowhead and right whales), Neobalaenidae (pygmy right whales), Eschirichtiidae (gray whales), and Balaenopteridae (rorquals). Phylogenetic relationships of the mysticeti order remain u ...
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Miocaperea
''Miocaperea'' is an extinct genus of pygmy right whale from the Miocene, Late Miocene Pisco Formation of Peru.''Miocaperea''
at Fossilworks.org
Its type species is ''Miocaperea pulchra''.


Evolution and significance

The discovery of ''Miocaperea'' is significant, because neobalaenines were previously unknown in the fossil record, except for an isolated Petrous part of the temporal bone, petrosal (ear bone) from late Miocene-aged deposits in Australia. A previous study placed the divergence date of Neobalaeninae from other mysticetes at about 23 million years,Sasaki T, Nikaido M, Hamilton H et al. (2005) Mitochondrial phylogenetics and evolution of mysticete whales. Systematic Biology 54(1): 77–90. and the age of ''Miocaperea'' lends credence to the notion of an origin ...
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Family (taxonomy)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Herpetocetus
''Herpetocetus'' is a genus of cetotheriid mysticete in the subfamily Herpetocetinae. Taxonomy There are four recognized species of ''Herpetocetus'': ''H. scaldiensis'', ''H. transatlanticus'',Whitmore, F.C., and L.G. Barnes. 2008. The Herpetocetinae, a new subfamily of extinct baleen whales (Mammalia, Cetacea, Cetotheriidae). In C.E. Ray, D.J. Bohaska, I.A. Koretsky, L.W. Ward, and L.G. Barnes (eds.). Geology and Paleontology of the Lee Creek Mine, North Carolina, IV. Virginia Museum of Natural History Special Publication 14:141–180. ''H. bramblei'', and ''H. morrowi''. An additional unnamed species from Japan has been referred to the '' nomen dubium'' '' Mizuhoptera sendaicus'' in the past, but the undiagnostic nature of the holotype of the latter means that it cannot be confidently referred to ''M. sendaicus''. Survival into the Pleistocene A recently discovered specimen of ''Herpetocetus'' from the Lower-Middle Pleistocene (Gelasian) Falor Formation of northern Califo ...
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Nannocetus
''Nannocetus'' is an extinct baleen whale belonging to the family Cetotheriidae. Description ''Nannocetus'' is a diminutive mysticete measuring 13 feet (4 meters) long. It is characterized by the ventral orientation (in posterior view) of the postglenoid process; postglenoid process twisted medially (in ventral view) relative to the anteroposterior axis of the skull; equal projection of the ventral and dorsal lobes of the tympanic than the dorsal lobe; deeper notch separating the two lobes of the tympanic; reniform morphology of the tympanic in ventral view; lip of the tympanic slightly inflated; sub-rectilinear medial edge of the involucrum with a step in its anterior third; anterior process of the petrosal sub-triangular; thin crista transversa of the petrosal; and pars cochlearis hemispherical. Taxonomy and classification The holotype is UCMP 26502. It was collected from the Late Miocene ( Messinian) Towsley Formation of Humphreys, Los Angeles County, California Cali ...
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