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Mycorrhizae
  A mycorrhiza (from Greek μύκης ', "fungus", and ῥίζα ', "root"; pl. mycorrhizae, mycorrhiza or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant's rhizosphere, its root system. Mycorrhizae play important roles in plant nutrition, soil biology, and soil chemistry. In a mycorrhizal association, the fungus colonizes the host plant's root tissues, either intracellularly as in arbuscular mycorrhizal fungi (AMF or AM), or extracellularly as in ectomycorrhizal fungi. The association is sometimes mutualistic. In particular species or in particular circumstances, mycorrhizae may have a parasitic association with host plants. Definition A mycorrhiza is a symbiotic association between a green plant and a fungus. The plant makes organic molecules such as sugars by photosynthesis and supplies them to the fungus, and the fungus supplies to the plant water and mineral nutrients, such as phosp ...
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Arbuscular Mycorrhiza
An arbuscular mycorrhiza (AM) (plural ''mycorrhizae'', a.k.a. ''endomycorrhiza'') is a type of mycorrhiza in which the symbiont fungus (''AM fungi'', or AMF) penetrates the cortical cells of the roots of a vascular plant forming arbuscules. (Not to be confused with ectomycorrhiza or ericoid mycorrhiza.) Arbuscular mycorrhizae are characterized by the formation of unique structures, arbuscules and vesicles, by Glomeromycota and Mucoromycota, sister clades of the more well-known and diverse dikaryan fungi (all three are together called "symbiomycota"). AM fungi help plants to capture nutrients such as phosphorus, sulfur, nitrogen and micronutrients from the soil. It is believed that the development of the arbuscular mycorrhizal symbiosis played a crucial role in the initial colonisation of land by plants and in the evolution of the vascular plants. It has been said that it is quicker to list the plants that do not form endomycorrhizae than those that do. This symbiosis is a ...
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Arbuscular Mycorrhizal Fungi
An arbuscular mycorrhiza (AM) (plural ''mycorrhizae'', a.k.a. ''endomycorrhiza'') is a type of mycorrhiza in which the symbiont fungus (''AM fungi'', or AMF) penetrates the cortical cells of the roots of a vascular plant forming arbuscules. (Not to be confused with ectomycorrhiza or ericoid mycorrhiza.) Arbuscular mycorrhizae are characterized by the formation of unique structures, arbuscules and vesicles, by Glomeromycota and Mucoromycota, sister clades of the more well-known and diverse dikaryan fungi (all three are together called "symbiomycota"). AM fungi help plants to capture nutrients such as phosphorus, sulfur, nitrogen and micronutrients from the soil. It is believed that the development of the arbuscular mycorrhizal symbiosis played a crucial role in the initial colonisation of land by plants and in the evolution of the vascular plants. It has been said that it is quicker to list the plants that do not form endomycorrhizae than those that do. This symbiosis is a ...
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Ectomycorrhiza
An ectomycorrhiza (from Greek ἐκτός ', "outside", μύκης ', "fungus", and ῥίζα ', "root"; pl. ectomycorrhizas or ectomycorrhizae, abbreviated EcM) is a form of symbiotic relationship that occurs between a fungal symbiont, or mycobiont, and the roots of various plant species. The mycobiont is often from the phyla Basidiomycota and Ascomycota, and more rarely from the Zygomycota. Ectomycorrhizas form on the roots of around 2% of plant species, usually woody plants, including species from the birch, dipterocarp, myrtle, beech, willow, pine and rose families. Research on ectomycorrhizas is increasingly important in areas such as ecosystem management and restoration, forestry and agriculture. Unlike other mycorrhizal relationships, such as arbuscular mycorrhiza and ericoid mycorrhiza, ectomycorrhizal fungi do not penetrate their host's cell walls. Instead they form an entirely intercellular interface known as the Hartig net, consisting of highly branched hyphae form ...
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Ectomycorrhiza
An ectomycorrhiza (from Greek ἐκτός ', "outside", μύκης ', "fungus", and ῥίζα ', "root"; pl. ectomycorrhizas or ectomycorrhizae, abbreviated EcM) is a form of symbiotic relationship that occurs between a fungal symbiont, or mycobiont, and the roots of various plant species. The mycobiont is often from the phyla Basidiomycota and Ascomycota, and more rarely from the Zygomycota. Ectomycorrhizas form on the roots of around 2% of plant species, usually woody plants, including species from the birch, dipterocarp, myrtle, beech, willow, pine and rose families. Research on ectomycorrhizas is increasingly important in areas such as ecosystem management and restoration, forestry and agriculture. Unlike other mycorrhizal relationships, such as arbuscular mycorrhiza and ericoid mycorrhiza, ectomycorrhizal fungi do not penetrate their host's cell walls. Instead they form an entirely intercellular interface known as the Hartig net, consisting of highly branched hyphae form ...
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Rhizosphere
The rhizosphere is the narrow region of soil or substrate that is directly influenced by root secretions and associated soil microorganisms known as the root microbiome. Soil pores in the rhizosphere can contain many bacteria and other microorganisms that feed on sloughed-off plant cells, termed ''rhizodeposition'', and the proteins and sugars released by roots, termed root exudates. This symbiosis leads to more complex interactions, influencing plant growth and competition for resources. Much of the nutrient cycling and disease suppression by antibiotics required by plants, occurs immediately adjacent to roots due to root exudates and metabolic products of symbiotic and pathogenic communities of microorganisms. The rhizosphere also provides space to produce allelochemicals to control neighbours and relatives. The ''rhizoplane'' refers to the root surface including its associated soil particles which closely interact with each other. The plant-soil feedback loop and other p ...
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Rhynie Chert
The Rhynie chert is a Lower Devonian sedimentary deposit exhibiting extraordinary fossil detail or completeness (a Lagerstätte). It is exposed near the village of Rhynie, Aberdeenshire, Scotland; a second unit, the Windyfield chert, is located some 700 m away. The Rhynie chert contains exceptionally preserved plant, fungus, lichen and animal material preserved in place by an overlying volcanic deposit. The bulk of the Devonian fossil bed consists of primitive plants (which had water-conducting cells and sporangia, but no true leaves), along with arthropods, lichens, algae and fungi. This fossil bed is remarkable for two reasons. First, the age of the site ( Pragian, Early Devonian, formed about ) places it at an early stage in the colonisation of land. Second, these cherts are famous for their exceptional state of ultrastructural preservation, with individual cell walls easily visible in polished specimens. Stomata have been counted and lignin remnants detected in the ...
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Flowering Plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils ar ...
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Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of the Mesozoic, Mesozoic Era and is named after the Jura Mountains, where limestone strata from the period were first identified. The start of the Jurassic was marked by the major Triassic–Jurassic extinction event, associated with the eruption of the Central Atlantic magmatic province, Central Atlantic Magmatic Province. The beginning of the Toarcian Stage started around 183 million years ago and is marked by an extinction event associated with widespread Anoxic event, oceanic anoxia, ocean acidification, and elevated temperatures likely caused by the eruption of the Karoo-Ferrar, Karoo-Ferrar large igneous provinces. The end of the Jurassic, however, has no clear boundary with the Cretaceous and i ...
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Aglaophyton
''Aglaophyton major'' (or more correctly ''Aglaophyton majus'') was the sporophyte generation of a diplohaplontic, pre-vascular, axial, free-sporing land plant of the Lower Devonian (Pragian stage, around ). It had anatomical features intermediate between those of the bryophytes and vascular plants or tracheophytes. ''A. major'' was first described by Kidston and Lang in 1920 as the new species ''Rhynia major''. The species is known only from the Rhynie chert in Aberdeenshire, Scotland, where it grew in the vicinity of a silica-rich hot spring, together with a number of associated vascular plants such as a smaller species ''Rhynia gwynne-vaughanii'' which may be interpreted as a representative of the ancestors of modern vascular plants and '' Asteroxylon mackei'', which was an ancestor of modern clubmosses (Lycopsida). Description The stems of ''Aglaophyton'' were round in cross-section, smooth, unornamented, and up to about 6mm in diameter. Kidston and Lang interpreted the pl ...
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New Phytologist
''New Phytologist'' is a peer-reviewed scientific journal published on behalf of the New Phytologist Foundation by Wiley-Blackwell. It was founded in 1902 by botanist Arthur Tansley, who served as editor until 1931. Topics covered ''New Phytologist'' covers all aspects of plant science, with topics ranging from intracellular processes through to global environmental change, including: * Physiology and development: intra/inter-cellular signalling, long-distance signalling, physiology, development, eco-devo - phenotypic plasticity, transport, biochemistry. * Environment: global change and Earth system functioning, environmental stress, ecophysiology, plant–soil interactions, heavy metals. * Interaction: multitrophic systems, mycorrhizas and pathogens, fungal genomics, nitrogen-fixing symbioses. * Evolution: molecular evolution, population genetics, mating systems, phylogenetics In biology, phylogenetics (; from Greek language, Greek wikt:φυλή, φυλή/wikt:φῦλο ...
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Timeline Of Plant Evolution
This article attempts to place key plant innovations in a geological context. It concerns itself only with novel adaptations and events that had a major ecological significance, not those that are of solely anthropological interest. The timeline displays a graphical representation of the adaptations; the text attempts to explain the nature and robustness of the evidence. Plant evolution is an aspect of the study of biological evolution, predominantly involving evolution of plants suited to live on land, greening of various land masses by the filling of their niches with land plants, and diversification of groups of land plants. Earliest plants In the strictest sense, the name ''plant'' refers to those land plants that form the clade Embryophyta, comprising the bryophytes and vascular plants. However, the clade Viridiplantae or green plants includes some other groups of photosynthetic eukaryotes, including green algae. It is widely believed that land plants evolved from a group ...
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Chenopodiaceae
Amaranthaceae is a family of flowering plants commonly known as the amaranth family, in reference to its type genus ''Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, making it the most species-rich lineage within its parent order, Caryophyllales. Description Vegetative characters Most species in the Amaranthaceae are annual or perennial herbs or subshrubs; others are shrubs; very few species are vines or trees. Some species are succulent. Many species have stems with thickened nodes. The wood of the perennial stem has a typical "anomalous" secondary growth; only in subfamily Polycnemoideae is secondary growth normal. The leaves are simple and mostly alternate, sometimes opposite. They never possess stipules. They are flat or terete, and their shape is extremely variable, with entire or toothed margins. In some species, the leaves are reduced to minute scales. In most cases, neither basal nor terminal aggrega ...
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