Modiolus Barbatus
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Modiolus Barbatus
''Modiolus barbatus'', the bearded horse mussel, is a species of "horse mussel", a marine (ocean), marine bivalve mollusc in the family (biology), family Mytilidae, the mussels. The fossil record of this species dates back to the Pliocene (age range: 3.6 to 2.588 million years ago). Description The shell of an adult ''Modiolus barbatus'' can be as long as .A. Lok, S. Acarli, S. Serdar, A. Kose and P. Goulletque GROWTH AND SURVIVAL RATES OF BEARDED HORSE MUSSEL (MODIOLUS BARBATUS LINNE, 1758) IN MERSIN BAY (TURKEY)/ref> The shape is elongated and ovoid or rhomboidal with a curved or obtuse dorsal margin. The outer surface of the shell is dark brown, while the inside of the valves is blue or slightly iridescent gray. These mussles are characterized by a ''periostracum'' with long, stiff, flat bristles. They are attached with strong ''byssus'' to the underground. Distribution and Habitat This species is present from the Mediterranean Sea to the Black Sea, on the southern and wester ...
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Pliocene
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58See the 2014 version of the ICS geologic time scale
million years ago. It is the second and most recent epoch of the Neogene Period in the . The Pliocene follows the Epoch and is followed by the Epoch. Prior to the 2009 ...
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Carl Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the modern system of naming organisms. He is known as the "father of modern taxonomy". Many of his writings were in Latin; his name is rendered in Latin as and, after his 1761 ennoblement, as . Linnaeus was born in Råshult, the countryside of Småland, in southern Sweden. He received most of his higher education at Uppsala University and began giving lectures in botany there in 1730. He lived abroad between 1735 and 1738, where he studied and also published the first edition of his ' in the Netherlands. He then returned to Sweden where he became professor of medicine and botany at Uppsala. In the 1740s, he was sent on several journeys through Sweden to find and classify plants and animals. In the 1750s and 1760s, he continued to collect an ...
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10th Edition Of Systema Naturae
The 10th edition of ''Systema Naturae'' is a book written by Swedish naturalist Carl Linnaeus and published in two volumes in 1758 and 1759, which marks the starting point of zoological nomenclature. In it, Linnaeus introduced binomial nomenclature for animals, something he had already done for plants in his 1753 publication of '' Species Plantarum''. Starting point Before 1758, most biological catalogues had used polynomial names for the taxa included, including earlier editions of ''Systema Naturae''. The first work to consistently apply binomial nomenclature across the animal kingdom was the 10th edition of ''Systema Naturae''. The International Commission on Zoological Nomenclature therefore chose 1 January 1758 as the "starting point" for zoological nomenclature, and asserted that the 10th edition of ''Systema Naturae'' was to be treated as if published on that date. Names published before that date are unavailable, even if they would otherwise satisfy the rules. The only ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Mussel
Mussel () is the common name used for members of several families of bivalve molluscs, from saltwater and Freshwater bivalve, freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other edible clams, which are often more or less rounded or oval. The word "mussel" is frequently used to mean the bivalves of the marine family Mytilidae, most of which live on exposed shores in the intertidal zone, attached by means of their strong Byssus, byssal threads ("beard") to a firm substrate. A few species (in the genus ''Bathymodiolus'') have colonised hydrothermal vents associated with deep ocean ridges. In most marine mussels the shell is longer than it is wide, being wedge-shaped or asymmetrical. The external colour of the shell is often dark blue, blackish, or brown, while the interior is silvery and somewhat nacreous. The common name "mussel" is also used for many freshwater bivalves, including the freshwater pearl mussels. F ...
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Marine (ocean)
The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the world ocean is conventionally divided."Ocean."
''Merriam-Webster.com Dictionary'', Merriam-Webster, https://www.merriam-webster.com/dictionary/ocean. Accessed March 14, 2021.
Separate names are used to identify five different areas of the ocean: (the largest), ,

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Bivalve
Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. They include the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. The shipworms bore into wood, clay, or stone and live inside these substances. The shell of a bivalve is composed of calc ...
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Mollusc
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8 taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastropods ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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Mytilidae
Mytilidae are a family of small to large marine and brackish-water bivalve molluscs in the order Mytilida. One of the genera, ''Limnoperna'', even inhabits freshwater environments. The order has only this one family which contains some 52 genera.Bouchet, P. (2014Mytilidae Rafinesque, 1815World Register of Marine Species Species in the family Mytilidae are found worldwide, but they are more abundant in colder seas, where they often form uninterrupted beds on rocky shores in the intertidal zone and the shallow subtidal. The subfamily Bathymodiolinae is found in deep-sea habitats. Mytilids include the well-known edible sea mussels. A common feature of the shells of mussels is an asymmetrical shell which has a thick, adherent periostracum. The animals attach themselves to a solid substrate using a byssus. A 2020 study of the phylogeny of Mytilidae recovered two main clades derived from an epifaunal ancestor, with subsequent lineages shifting to other lifestyles, and correlat ...
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Periostracum
The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and bivalves, but it is also found in cephalopods such as ''Allonautilus scrobiculatus''. The periostracum is an integral part of the shell, and it forms as the shell forms, along with the other shell layers. The periostracum is used to protect the organism from corrosion. The periostracum is visible as the outer layer of the shell of many molluscan species from terrestrial, freshwater, and marine habitats, and may be seen in land snails, river mussels, and other kinds of freshwater bivalves, as well as in many kinds of marine shelled molluscs. The word ''periostracum'' means "around the shell", meaning that the periostracum is wrapped around what is usually the more calcareous part of the shell. Technically, the calcareous part of the shel ...
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Byssus
A byssus () is a bundle of filaments secreted by many species of bivalve mollusc that function to attach the mollusc to a solid surface. Species from several families of clams have a byssus, including pen shells (Pinnidae), true mussels (Mytilidae), and Dreissenidae. Filaments Byssus filaments are created by certain kinds of marine and freshwater bivalve mollusks, which use the byssus to attach themselves to rocks, substrates, or seabeds. In edible mussels, the inedible byssus is commonly known as the "beard", and is removed before cooking. Many species of mussels secrete byssus threads to anchor themselves to surfaces, with families including the Arcidae, Mytilidae, Anomiidae, Pinnidae, Pectinidae, Dreissenidae, and Unionidae. Mechanics The byssus, or byssal complex, is composed of multiple extracellular collagenous threads that are placed radially by the mussel from a central stem. Each thread is composed of three regions: a corrugated proximal region close to the mussel b ...
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