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Micoletzkya
''Micoletzkya'' is a genus of predatory bark-beetle-associated nematodes in the family Diplogastridae. The genus was named for the Austrian nematologist Heinrich Micoletzky. As of 2013 it consists of 25 described species. Ecology and life history Nematodes of ''Micoletzkya'' live in symbiosis with bark beetles.Rühm, W. 1956. Die Nematoden der Ipiden. Parasitologische Schriftenreihe No. 6: 1–437. The hosts include economically important species across North America and Europe such as the mountain pine beetle ''Dendroctonus ponderosae'' and European spruce bark beetle ''Ips typographus''. The life history of the nematodes is closely linked to that of their hosts. Infective ( dauer) larvae of ''Micoletzkya'' attach themselves onto the body surface of a beetle for transmission and resume their development in host breeding galleries. In the bark beetle galleries, nematodes go through a few free-living generations, presumably feeding on beetle-associated microorganisms and other n ...
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Diplogastridae
Diplogastridae, formerly Diplogasteridae, are a family of nematodes (roundworms) known from a wide range of habitats, often in commensal or parasitic associations with insects. Description Diplogastrid nematodes are characterized by a distinct "two-lobed" pharynx (hence their name from the Greek διπλόος = "double" and γαστήρ = "stomach"), the second ( posterior) lobe being composed mostly of glandular tissue. Most known species also have at least one tooth, which has presumably allowed them to access many new food sources compared with the related nematodes of Rhabditidae (including '' Caenorhabditis elegans''), most species of which feed on bacteria. Several diplogastrid species also have a polyphenism in their mouthparts, allowing resource specialization within species. The wide array of feeding modes in the Diplogastridae is reflected by the relatively high diversity and complexity of their mouth structures, which show accelerated rates of evolution in comparis ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Nematode
The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-Parasitism, parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broad range of environments. Less formally, they are categorized as Helminths, but are taxonomically classified along with Arthropod, arthropods, Tardigrade, tardigrades and other moulting animalia, animals in the clade Ecdysozoa, and unlike platyhelminthe, flatworms, have tubular digestion, digestive systems with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum. Nematode species can be difficult to distinguish from one another. Consequently, estimates of the number of nematode species described to date vary by author and may change rapidly over ...
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Chromadorea
The Chromadorea are a class of the roundworm phylum, Nematoda. They contain a single subclass (Chromadoria) and several orders. With such a redundant arrangement, the Chromadoria are liable to be divided if the orders are found to form several clades, or abandoned if they are found to constitute a single radiation. Formerly, they were treated as a subclass in the paraphyletic "Adenophorea" assemblage, which has been mostly abandoned by modern authors. It is also suspected that the Chromadorea may not be monophyletic as delimited here; at least the Monhysterida seem to be a distinct and far more ancient lineage than the rest. Members of this class' bodies usually have annules, their amphids elaborate and spiral, and they all have three esophageal glands. They usually live in marine sediments, although they can live elsewhere. They have a more sophisticated pharynx than most roundworms. Members of this class can be identified by the presence of eight conserved signature indels (C ...
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Rhabditida
Rhabditida is an order of free-living, zooparasitic, and phytoparasitic microbivorous nematodes living in soil. The Cephalobidae, Panagrolaimidae, Steinernematidae, and Strongyloididae seem to be closer to the Tylenchia, regardless of whether these are merged with the Rhabditia or not.Tree of Life Web Project (2002b)Nematoda Version of 2002-JAN-01. Retrieved 2008-NOV-02. Families Rhabditida * Myolaimina **''Incertae sedis'' *** Myolaimoidea ****Myolaimidae *Rhabditina ** Bunonematomorpha *** Bunonematoidea ****Bunonematidae ** Diplogasteromorpha *** Cylindrocorporoidea ****Cylindrocorporidae *** Diplogasteroidea **** Cephalobiidae ****Diplogasteridae ****Diplogasteroididae ** Rhabditomorpha *** Mesorhabditoidea ****Peloderidae *** Rhabditoidea ****Rhabditidae * Spirurina ** Ascaridomorpha *** Ascaridoidea **** Acanthocheilidae ****Anisakidae ****Ascarididae ****Heterocheilidae **** Raphidascarididae *** Cosmocercoidea ****Atractidae ****Kathlaniidae *** Seuratoidea **** Cuc ...
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Symbiosis
Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasitic. The organisms, each termed a symbiont, must be of different species. In 1879, Heinrich Anton de Bary defined it as "the living together of unlike organisms". The term was subject to a century-long debate about whether it should specifically denote mutualism, as in lichens. Biologists have now abandoned that restriction. Symbiosis can be obligatory, which means that one or more of the symbionts depend on each other for survival, or facultative (optional), when they can generally live independently. Symbiosis is also classified by physical attachment. When symbionts form a single body it is called conjunctive symbiosis, while all other arrangements are called disjunctive symbiosis."symbiosis." Dorland's Illustrated Medical Dictionary. ...
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Bark Beetles
A bark beetle is the common name for the subfamily of beetles Scolytinae. Previously, this was considered a distinct family (Scolytidae), but is now understood to be a specialized clade of the "true weevil" family (Curculionidae). Although the term "bark beetle" refers to the fact that many species feed in the inner bark (phloem) layer of trees, the subfamily also has many species with other lifestyles, including some that bore into wood, feed in fruit and seeds, or tunnel into herbaceous plants. Well-known species are members of the type genus ''Scolytus'', namely the European elm bark beetle ''S. multistriatus'' and the large elm bark beetle ''S. scolytus'', which like the American elm bark beetle ''Hylurgopinus rufipes'', transmit Dutch elm disease fungi (''Ophiostoma''). The mountain pine beetle ''Dendroctonus ponderosae'', southern pine beetle ''Dendroctonus frontalis'', and their near relatives are major pests of conifer forests in North America. A similarly aggressive spec ...
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Mountain Pine Beetle
The mountain pine beetle (''Dendroctonus ponderosae'') is a species of bark beetle native to the forests of western North America from Mexico to central British Columbia. It has a hard black exoskeleton, and measures approximately , about the size of a grain of rice. In western North America, a recent outbreak of the mountain pine beetle and its microbial associates has affected wide areas of lodgepole pine forest, including more than of forest in British Columbia. The outbreak in the Rocky Mountain National Park in Colorado began in 1996 and has caused the destruction of millions of acres/hectares of ponderosa and lodgepole pine trees. At the peak of the outbreak in 2009, over were affected. The outbreak then declined due to better environmental conditions and the fact that many vulnerable trees had been already destroyed. Mountain pine beetles inhabit ponderosa, whitebark, lodgepole, Scots, jack, limber, Rocky Mountain bristlecone, and Great Basin bristlecone pine tre ...
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European Spruce Bark Beetle
The European spruce bark beetle (''Ips typographus''), is a species of beetle in the weevil subfamily Scolytinae, the bark beetles, and is found from Europe to Asia Minor and some parts of Africa. Biology of the species Morphology Adults are usually long, cylindrical and robust, black or brownish-black. Elytral declivity is slightly shiny, with 4 teeth on each margin side. The third tooth is the biggest and club like on its top. The egg is yellowish-white. The larva is white and legless. The pupa is also white. Life cycle and interactions Bark beetles are so named because they reproduce in the inner bark, living and dead phloem tissues, of trees. Adult beetles hibernate in forest litter and host trees when environmental conditions are not favorable for reproduction. When conditions are right, they travel up to half a mile in search of a vulnerable host. Once the host is located, the adult burrows through the weakened bark in order to build tunnels where they can mate and l ...
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Dauer Larvae
Dauer (German "''die Dauer''", English "the enduring", "the duration" in the meaning of "a length of time",) describes an alternative developmental stage of nematode worms, particularly rhabditids including ''Caenorhabditis elegans'', whereby the larva goes into a type of stasis and can survive harsh conditions. Since the entrance of the dauer stage is dependent on environmental cues, it represents a classic and well studied example of polyphenism. The dauer state is given other names in the various types of nematodes such as ‘ diapause’ or ‘hypobiosis’, but since the ''C. elegans'' nematode has become the most studied nematode, the term ‘dauer stage’ or 'dauer larvae' is becoming universally recognised when referring to this state in other free-living nematodes. The dauer stage is also considered to be equivalent to the infective stage of parasitic nematode larvae. As E. Maupas first proposed in 1899-1900, all nematodes have five stages separated by four moults. Und ...
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Phenotypes
In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological properties, its behavior, and the products of behavior. An organism's phenotype results from two basic factors: the expression of an organism's genetic code, or its genotype, and the influence of environmental factors. Both factors may interact, further affecting phenotype. When two or more clearly different phenotypes exist in the same population of a species, the species is called polymorphic. A well-documented example of polymorphism is Labrador Retriever coloring; while the coat color depends on many genes, it is clearly seen in the environment as yellow, black, and brown. Richard Dawkins in 1978 and then again in his 1982 book ''The Extended Phenotype'' suggested that one can regard bird nests and other built structures such as cad ...
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Polyphenism
A polyphenic trait is a trait for which multiple, discrete phenotypes can arise from a single genotype as a result of differing environmental conditions. It is therefore a special case of phenotypic plasticity. There are several types of polyphenism in animals, from having sex determined by the environment to the castes of honey bees and other social insects. Some polyphenisms are seasonal, as in some butterflies which have different patterns during the year, and some Arctic animals like the snowshoe hare and Arctic fox, which are white in winter. Other animals have predator-induced or resource polyphenisms, allowing them to exploit variations in their environment. Some nematode worms can develop either into adults or into resting dauer larvae according to resource availability. Definition upright=1.2, Polyphenism in termites A : Primary king B : Primary queen C : Secondary queen D : Tertiary queen E : Soldiers F : Worker A polyphenism is the occurr ...
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