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Metriocnemus Knabi
''Metriocnemus knabi'', the pitcher plant midge, is an inquiline invertebrate found only in the phytotelma of the purple pitcher plant, ''Sarracenia purpurea''. In this microcommunity of bacteria, rotifers, protozoa, and other dipteran larva like ''Wyeomyia smithii ''Wyeomyia smithii'', the pitcher plant mosquito, is an inquiline mosquito that completes its pre-adult life cycle in the phytotelma of—that is, the water contained by—the purple pitcher plant, ''Sarracenia purpurea''. In this microcommunity ...'', ''M. knabi'' specializes by feeding mostly on the carcasses of drowned insects captured by the plant that collect at the bottom of the pitcher. It is found in the eastern United States, as far north as Maine and south to North Carolina. References Chironomidae Sarracenia purpurea inquilines Diptera of North America Insects described in 1904 Taxa named by Daniel William Coquillett {{Chironomidae-stub ...
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Daniel William Coquillett
Daniel William Coquillett (23 January 1856, Pleasant Valley, Ill. – 7 July 1911 Atlantic City, New Jersey) was an American entomologist who specialised in Diptera. He wrote a revision of the dipterous family Therevidae and many other scientific papers in which he described many new species and genera of Diptera. Coquillett was also the first to attempt fumigation with hydrocyanic acid as a means for controlling citrus scale insects. He experimented in the Wolfskill orange groves where he was supported by the foreman and later quarantine entomologist Alexander Craw Alexander Craw (3 August 1850 – 28 June 1908) was a pioneer American economic entomologist. He was the first American entomologist to work in quarantine protection against foreign pests arriving by ship to San Francisco, California. Along with ... in 1888–89. References External linksArchiveDigitised Coquillett, D. W. ''Report on the locusts of the San Joaquin valley, Cal.'' Anaheim, Calif.Date 1886ArchiveD ...
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Inquiline
In zoology, an inquiline (from Latin ''inquilinus'', "lodger" or "tenant") is an animal that lives commensally in the nest, burrow, or dwelling place of an animal of another species. For example, some organisms such as insects may live in the homes of gophers or the garages of humans and feed on debris, fungi, roots, etc. The most widely distributed types of inquiline are those found in association with the nests of social insects, especially ants and termites – a single colony may support dozens of different inquiline species. The distinctions between parasites, social parasites, and inquilines are subtle, and many species may fulfill the criteria for more than one of these, as inquilines do exhibit many of the same characteristics as parasites. However, parasites are specifically ''not'' inquilines, because by definition they have a deleterious effect on the host species, while inquilines have not been confirmed to do so. In the specific case of termites, the term "'' ...
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Phytotelma
Phytotelma (plural phytotelmata) is a small water-filled cavity in a terrestrial plant. The water accumulated within these plants may serve as the habitat for associated fauna and flora. A rich literature in German summarised by Thienemann (1954) developed many aspects of phytotelm biology. Reviews of the subject by Kitching (1971) and Maguire (1971) introduced the concept of phytotelmata to English-speaking readers. A multi-authored book edited by Frank and Lounibos (1983)Frank, J.H. and Lounibos, L.P. (1983) ''Phytotelmata: Terrestrial plants as hosts for aquatic insect communities'', Plexus Press. dealt in 11 chapters with classification of phytotelmata, and with phytotelmata provided by bamboo internodes, banana leaf axils, bromeliad leaf axils, ''Nepenthes'' pitchers, ''Sarracenia'' pitchers, tree holes, and ''Heliconia'' flower bracts and leaf rolls. A classification of phytotelmata by Kitching (2000) recognizes five principal types: bromeliad tanks, certain carnivor ...
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Sarracenia Purpurea
''Sarracenia purpurea'', the purple pitcher plant, northern pitcher plant, turtle socks, or side-saddle flower, is a carnivorous plant in the family Sarraceniaceae. Description Like other species of ''Sarracenia'', ''S. purpurea'' obtains most of its nutrients through prey capture. However, prey acquisition is said to be inefficient, with less than 1% of the visiting prey captured within the pitcher. Even so, anecdotal evidence by growers often shows that pitchers quickly fill up with prey during the warm summer months. Prey fall into the pitcher and drown in the rainwater that collects in the base of each leaf. Prey items, such as flies, ants, spiders, and even moths or hornets, are then digested by an invertebrate community, made up mostly by the mosquito ''Wyeomyia smithii'' and the midge ''Metriocnemus knabi''. The relationship between ''W. smithii'' and ''S. purpurea'' is an example of commensalism. ''S. purpurea'' also traps juvenile spotted salamanders with enough reg ...
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Rotifer
The rotifers (, from the Latin , "wheel", and , "bearing"), commonly called wheel animals or wheel animalcules, make up a phylum (Rotifera ) of microscopic and near-microscopic pseudocoelomate animals. They were first described by Rev. John Harris in 1696, and other forms were described by Antonie van Leeuwenhoek in 1703. Most rotifers are around long (although their size can range from to over ), and are common in freshwater environments throughout the world with a few saltwater species. Some rotifers are free swimming and truly planktonic, others move by inchworming along a substrate, and some are sessile, living inside tubes or gelatinous holdfasts that are attached to a substrate. About 25 species are colonial (e.g., '' Sinantherina semibullata''), either sessile or planktonic. Rotifers are an important part of the freshwater zooplankton, being a major foodsource and with many species also contributing to the decomposition of soil organic matter. Most species of the r ...
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Protozoa
Protozoa (singular: protozoan or protozoon; alternative plural: protozoans) are a group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic tissues and debris. Historically, protozoans were regarded as "one-celled animals", because they often possess animal-like behaviours, such as motility and predation, and lack a cell wall, as found in plants and many algae. When first introduced by Georg Goldfuss (originally spelled Goldfuß) in 1818, the taxon Protozoa was erected as a class within the Animalia, with the word 'protozoa' meaning "first animals". In later classification schemes it was elevated to a variety of higher ranks, including phylum, subkingdom and kingdom, and sometimes included within Protoctista or Protista. The approach of classifying Protozoa within the context of Animalia was widespread in the 19th and early 20th century, but not universal. By the 1970s, it became usual to require th ...
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Wyeomyia Smithii
''Wyeomyia smithii'', the pitcher plant mosquito, is an inquiline mosquito that completes its pre-adult life cycle in the phytotelma of—that is, the water contained by—the purple pitcher plant, ''Sarracenia purpurea''. In this microcommunity of bacteria, rotifers, protozoa, and midges, ''W. smithii'' is the top-level predator; its presence determines the bacterial species diversity within the pitcher. ''W. smithii'' is not a pest mosquito in general. The northern US population does not consume blood at all, while the southern US populations only consume blood after laying an initial egg batch; even then they appear disinterested in feeding. In fact, it is the only known mosquito to have both obligatory biting and non-biting populations in the same species. Life cycle Description of the life cycle The life cycle of ''Wyeomyia smithii'' begins in either late spring or early fall when the adult female lays her eggs in the phytolema of a purple pitcher plant. The eggs then ge ...
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Chironomidae
The Chironomidae (informally known as chironomids, nonbiting midges, or lake flies) comprise a family of nematoceran flies with a global distribution. They are closely related to the Ceratopogonidae, Simuliidae, and Thaumaleidae. Many species superficially resemble mosquitoes, but they lack the wing scales and elongated mouthparts of the Culicidae. The name Chironomidae stems from the Ancient Greek word ''kheironómos'', "a pantomimist". Common names and biodiversity This is a large taxon of insects; some estimates of the species numbers suggest well over 10,000 world-wide. Males are easily recognized by their plumose antennae. Adults are known by a variety of vague and inconsistent common names, largely by confusion with other insects. For example, chironomids are known as "lake flies" in parts of Canada and Lake Winnebago, Wisconsin, but "bay flies" in the areas near the bay of Green Bay, Wisconsin. They are called "sand flies", "muckleheads", "muffleheads", "Canadian so ...
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Sarracenia Purpurea Inquilines
''Sarracenia'' ( or ) is a genus comprising 8 to 11 species of North American pitcher plants, commonly called trumpet pitchers. The genus belongs to the family Sarraceniaceae, which also contain the closely allied genera '' Darlingtonia'' and ''Heliamphora''. ''Sarracenia'' is a genus of carnivorous plants indigenous to the eastern seaboard of the United States, Texas, the Great Lakes area and southeastern Canada, with most species occurring only in the south-east United States (only '' S. purpurea'' occurs in cold-temperate regions). The plant's leaves have evolved into a funnel or pitcher shape in order to trap insects. The plant attracts its insect prey with secretions from extrafloral nectaries on the lip of the pitcher leaves, as well as a combination of the leaves' color and scent. Slippery footing at the pitcher's rim, causes insects to fall inside, where they die and are digested by the plant with proteases and other enzymes. Description ''Sarracenia'' are he ...
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Diptera Of North America
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies and others, although only about 125,000 species have been described. Flies have a mobile head, with a pair of large compound eyes, and mouthparts designed for piercing and sucking (mosquitoes, black flies and robber flies), or for lapping and sucking in the other groups. Their wing arrangement gives them great maneuverability in flight, and claws and pads on their feet enable them to cling to smooth surfaces. Flies undergo complete metamorphosis; the eggs are often laid on the la ...
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Insects Described In 1904
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; In: potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within. Nearly all insects hatch from eggs. Insect ...
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