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Melezitose
Melezitose, also spelled melicitose, is a nonreducing trisaccharide sugar that is produced by many plant sap eating insects, including aphids such as ''Cinara pilicornis'', by an enzyme reaction. This is beneficial to the insects, as it reduces the stress of osmosis by reducing their own water potential. The melezitose is part of the Honeydew (secretion), honeydew which acts as an attractant for ants and also as a food for bees. This is useful to the aphids as they have a symbiotic relationship with ants. Melezitose can be partially hydrolysis, hydrolyzed to glucose and turanose the latter of which is an isomer of sucrose. References

{{Carbohydrates Trisaccharides ...
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Trisaccharide
''Trisaccharides'' are oligosaccharides composed of three monosaccharides with two glycosidic bonds connecting them. Similar to the disaccharides, each glycosidic bond can be formed between any hydroxyl group on the component monosaccharides. Even if all three component sugars are the same (e.g., glucose Glucose is a simple sugar with the molecular formula . Glucose is overall the most abundant monosaccharide, a subcategory of carbohydrates. Glucose is mainly made by plants and most algae during photosynthesis from water and carbon dioxide, u ...), different bond combinations (regiochemistry) and stereochemistry (alpha- or beta-) result in trisaccharides that are diastereoisomers with different chemical and physical properties. Examples References 1) Lehninger Principles of Biochemistry by Albert L. Lehninger, David L. Nelson, and Michael M. Cox. 2) Biochemistry by Lubert Stryer. External links * {{organic-compound-stub ...
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Sugar
Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose, fructose, and galactose. Compound sugars, also called disaccharides or double sugars, are molecules made of two bonded monosaccharides; common examples are sucrose (glucose + fructose), lactose (glucose + galactose), and maltose (two molecules of glucose). White sugar is a refined form of sucrose. In the body, compound sugars are hydrolysed into simple sugars. Longer chains of monosaccharides (>2) are not regarded as sugars, and are called oligosaccharides or polysaccharides. Starch is a glucose polymer found in plants, the most abundant source of energy in human food. Some other chemical substances, such as glycerol and sugar alcohols, may have a sweet taste, but are not classified as sugar. Sugars are found in the tissues of most plants. Honey and fruits are abundant natural sources of simple sugars. Suc ...
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Plant Sap
Sap is a fluid transported in xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a separate substance, separately produced, and with different components and functions. Insect honeydew is called sap, particularly when it falls from trees, but is only the remains of eaten sap and other plant parts. Types of sap Saps may be broadly divided into two types: xylem sap and phloem sap. Xylem sap Xylem sap (pronounced ) consists primarily of a watery solution of hormones, mineral elements and other nutrients. Transport of sap in xylem is characterized by movement from the roots toward the leaves. Over the past century, there has been some controversy regarding the mechanism of xylem sap transport; today, most plant scientists agree that the cohesion-tension theory best explains this process, but multiforce theories that hy ...
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Aphid
Aphids are small sap-sucking insects and members of the superfamily Aphidoidea. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white woolly aphids. A typical life cycle involves flightless females giving live birth to female nymphs—who may also be already pregnant, an adaptation scientists call telescoping generations—without the involvement of males. Maturing rapidly, females breed profusely so that the number of these insects multiplies quickly. Winged females may develop later in the season, allowing the insects to colonize new plants. In temperate regions, a phase of sexual reproduction occurs in the autumn, with the insects often overwintering as eggs. The life cycle of some species involves an alternation between two species of host plants, for example between an annual crop and a woody plant. Some species feed on only one type of plant, while others are generalists, coloni ...
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Cinara Pilicornis
''Cinara pilicornis'', the spruce shoot aphid or brown spruce shoot aphid, is an aphid species in the genus ''Cinara'' found on Norway spruce (''Picea abies'') and Sitka spruce (''Picea sitchensis'').An Entomophthora Species on Cinara pilicornis (Hartig) (Hemiptera: Aphididae). T. L. Edwards, The Irish Naturalists' Journal, Jan. 1981, Vol. 20, No. 5, pages 204-206jstor It is a quite large aphid species with a plump, dull brown body. It seems to have little effect on the tree. It is a European species but it has also been reported in spruce forests in New Zealand, together with the spruce aphid ('' Elatobium abietinum''). ''C. pilicornis'', which is attended by the honeydew-collecting ants ''Formica polyctena'', is seldom attacked by the parasitoid wasp '' Pauesia pini''. It is also a host for ''Entomophthora'' fungi. ''C. pilicornis'' produces the trisaccharide melezitose. Citronellol, cis–trans-nepetalactone and cis–trans-nepetalactol Nepetalactol is an iridoid. It is pr ...
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Insects
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; In: potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within. Nearly all insects hatch from eggs. Inse ...
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Osmosis
Osmosis (, ) is the spontaneous net movement or diffusion of solvent molecules through a selectively-permeable membrane from a region of high water potential (region of lower solute concentration) to a region of low water potential (region of higher solute concentration), in the direction that tends to equalize the solute concentrations on the two sides. It may also be used to describe a physical process in which any solvent moves across a selectively permeable membrane (permeable to the solvent, but not the solute) separating two solutions of different concentrations. Osmosis can be made to do work. Osmotic pressure is defined as the external pressure required to be applied so that there is no net movement of solvent across the membrane. Osmotic pressure is a colligative property, meaning that the osmotic pressure depends on the molar concentration of the solute but not on its identity. Osmosis is a vital process in biological systems, as biological membranes are semiperm ...
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Honeydew (secretion)
Honeydew is a sugar-rich sticky liquid, secreted by aphids and some scale insects as they feed on plant sap. When their mouthpart penetrates the phloem, the sugary, high-pressure liquid is forced out of the anus of the aphid. Honeydew is particularly common as a secretion in hemipteran insects and is often the basis for trophobiosis. Some caterpillars of Lycaenidae butterflies and some moths also produce honeydew. Honeydew producing insects, like cicadas, pierce phloem ducts to access the sugar rich sap. The sap continues to bleed after the insects have moved on, leaving a white sugar crust called manna. Ants may collect, or "milk", honeydew directly from aphids and other honeydew producers, which benefit from their presence due to their driving away predators such as lady beetles or parasitic wasps—see ''Crematogaster peringueyi''. Animals and plants in a mutually symbiotic arrangement with ants are called Myrmecophiles. In Madagascar, some gecko species in the genera ''Ph ...
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Ants
Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22,000 species have been classified. They are easily identified by their geniculate (elbowed) antennae and the distinctive node-like structure that forms their slender waists. Ants form colonies that range in size from a few dozen predatory individuals living in small natural cavities to highly organised colonies that may occupy large territories and consist of millions of individuals. Larger colonies consist of various castes of sterile, wingless females, most of which are workers (ergates), as well as soldiers (dinergates) and other specialised groups. Nearly all ant colonies also have some fertile males called "drones" and one or more fertile females called "queens" (gynes). The colonies are described as superorganisms because the ants appea ...
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Hydrolysis
Hydrolysis (; ) is any chemical reaction in which a molecule of water breaks one or more chemical bonds. The term is used broadly for substitution reaction, substitution, elimination reaction, elimination, and solvation reactions in which water is the nucleophile. Biological hydrolysis is the cleavage of biomolecules where a water molecule is consumed to effect the separation of a larger molecule into component parts. When a carbohydrate is broken into its component sugar molecules by hydrolysis (e.g., sucrose being broken down into glucose and fructose), this is recognized as saccharification. Hydrolysis reactions can be the reverse of a condensation reaction in which two molecules join into a larger one and eject a water molecule. Thus hydrolysis adds water to break down, whereas condensation builds up by removing water. Types Usually hydrolysis is a chemical process in which a molecule of water is added to a substance. Sometimes this addition causes both the substance and w ...
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Glucose
Glucose is a simple sugar with the molecular formula . Glucose is overall the most abundant monosaccharide, a subcategory of carbohydrates. Glucose is mainly made by plants and most algae during photosynthesis from water and carbon dioxide, using energy from sunlight, where it is used to make cellulose in cell walls, the most abundant carbohydrate in the world. In energy metabolism, glucose is the most important source of energy in all organisms. Glucose for metabolism is stored as a polymer, in plants mainly as starch and amylopectin, and in animals as glycogen. Glucose circulates in the blood of animals as blood sugar. The naturally occurring form of glucose is -glucose, while -glucose is produced synthetically in comparatively small amounts and is less biologically active. Glucose is a monosaccharide containing six carbon atoms and an aldehyde group, and is therefore an aldohexose. The glucose molecule can exist in an open-chain (acyclic) as well as ring (cyclic) form. Gluco ...
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Turanose
Turanose is a reducing disaccharide. The -isomer is naturally occurring. Its systematic name is α--glucopyranosyl-(1→3)-α--fructofuranose. It is an analog of sucrose not metabolized by higher plants, but rather acquired through the action of sucrose transporters for intracellular carbohydrate signaling. In addition to its involvement in signal transduction, -(+)-turanose can also be used as a carbon source by many organisms including numerous species of bacteria and fungi.D-Turanose
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Sigma-Aldrich Sigma-Aldrich (formally MilliporeSigma) is an American chemical, life science, and biotechnology company that is owned by the German chemical conglomerate Merck Group. Sigma-Aldrich wa ...
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