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Maroccosuchus
''Maroccosuchus zennaroi'' is an extinct gavialoid crocodylian from the Early Eocene of Morocco, traditionally regarded as a member of the subfamily Tomistominae. Below is a cladogram based on morphological studies comparing skeletal features that shows ''Maroccosuchus'' as a member of Tomistominae, related to the false gharial: Based on morphological studies of extinct taxa, the tomistomines were long thought to be classified as crocodiles and not closely related to gavialoids. However, recent molecular studies using DNA sequencing have consistently indicated that the false gharial (''Tomistoma'') (and by inference other related extinct forms in Tomistominae) actually belong to Gavialoidea (and Gavialidae). Below is a cladogram from a 2018 tip dating study by Lee & Yates simultaneously using morphological, molecular (DNA sequencing), and stratigraphic (fossil age) data that shows ''Maroccosuchus'' as an early-diverging gavialoid, more basal than the last common ancestor to ...
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Tomistominae
Tomistominae is a subfamily of crocodylians that includes one living species, the false gharial. Many more extinct species are known, extending the range of the subfamily back to the Eocene epoch. In contrast to the false gharial, which is a freshwater species that lives only in southeast Asia, extinct tomistomines had a global distribution and lived in estuaries and along coastlines. The classification of tomistomines among Crocodylia has been in flux; while traditionally thought to be within Crocodyloidea, molecular evidence indicates that they are more closely related to true gharials as members of Gavialoidea. Description Tomistomines have narrow or longirostrine snouts like gharials. The living false gharial lives in fresh water and uses its long snout and sharp teeth to catch fish, although true gharials are more adapted toward piscivory, or fish-eating. Despite the similarity with gharials, the shapes of bones in tomistomine skulls link them with crocodiles. For example, ...
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Gavialidae
Gavialidae is a family (biology), family of large semiaquatic crocodilians with elongated, narrow snouts. Gavialidae consists of two extant taxon, living species, the gharial (''Gavialis gangeticus'') and the false gharial (''Tomistoma schlegelii''), both occurring in Asia. Many extinct members are known from a broader range, including the recently extinct ''Hanyusuchus''. Gavialids are generally regarded as lacking the jaw strength to capture the large mammalian prey favoured by crocodiles and alligators of similar size so their thin snout is best used to catch fish, however the false gharial has been found to have a Generalist and specialist species, generalist diet with mature adults preying upon larger vertebrates, such as Ungulate, ungulates. Taxonomy The family Gavialidae was proposed by Arthur Adams (zoologist), Arthur Adams in 1854 for reptiles with a very long and slender muzzle, webbed feet and nearly equal teeth. It is currently recognized as a crown group, meaning that ...
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Gavialoid
Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomistoma schlegelii'' are alive today, with ''Hanyusuchus'' having become extinct in the last few centuries. Extinct South American gavialoids likely dispersed in the mid Tertiary from Africa and Asia. Fossil remains of the Puerto Rican gavialoid '' Aktiogavialis puertorisensis'' were discovered in a cave located in San Sebastián, Puerto Rico and dated to the Oligocene. This individual is thought to have crossed the Atlantic coming from Africa, indicating that this species was able to withstand saltwater. Classification Gavialoidea is cladistically defined as ''Gavialis gangeticus'' (the gharial) and all crocodylians closer to it than to ''Alligator mississippiensis'' (the American alligator) or ''Crocodylus niloticus'' (the Nile crocodile). This is a stem-bas ...
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Gavialoidea
Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomistoma schlegelii'' are alive today, with ''Hanyusuchus'' having become extinct in the last few centuries. Extinct South American gavialoids likely dispersed in the mid Tertiary from Africa and Asia. Fossil remains of the Puerto Rican gavialoid '' Aktiogavialis puertorisensis'' were discovered in a cave located in San Sebastián, Puerto Rico and dated to the Oligocene. This individual is thought to have crossed the Atlantic coming from Africa, indicating that this species was able to withstand saltwater. Classification Gavialoidea is cladistically defined as ''Gavialis gangeticus'' (the gharial) and all crocodylians closer to it than to ''Alligator mississippiensis'' (the American alligator) or ''Crocodylus niloticus'' (the Nile crocodile). This is a stem-bas ...
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Dollosuchoides
''Dollosuchoides'', colloquially known as the Crocodile of Maransart, is an extinct monospecific genus of gavialoid crocodilian, traditionally regarded as a member of the subfamily Tomistominae. Fossils have been found in the Brussel Formation of Maransart, Belgium and date back to the middle Eocene. The holotype, IRScNB 482, was discovered in 1915 and it was prepared during 1926–1927 by M. Hubert, J. Mehschaert and M. Jean de Kleermaeker, and also in 1927, Louis Dollo had the holotype put on display in the Museum of Natural Sciences and he intended to describe the specimen but he died in 1931 before he was able to describe it and the specimen was eventually referred to '' Dollosuchus'' by Swinton (1937)Swinton, W. E. (1937)The Crocodile of Maransart (Dollosuchus dixoni [Owen]).''Mémoire'' 80: 3–46 until it was moved to its own genus by Brochu (2007). It is currently housed in the Gand Museum in Belgium. Phylogeny Below is a cladogram based morphological studies c ...
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Tomistoma Cairense
''Tomistoma cairense'' is an extinct species of gavialoid crocodilian from the Lutetian stage of the Eocene era. It lived in North East Africa, especially Egypt. Remains of ''T. cairense'' have been found in the Mokattam Formation, in Mokattam, Egypt. ''Tomistoma cairense'' did not have a Maxilla process within their lacrimal gland, whereas all extant (living) crocodilians do. Below is a cladogram based morphological studies comparing skeletal features that shows ''Tomistoma cairense'' as a member of Tomistominae, related to the false gharial: Based on morphological studies of extinct taxa, the tomistomines (including the living false gharial) were long thought to be classified as crocodiles and not closely related to gavialoids. However, recent molecular studies using DNA sequencing have consistently indicated that the false gharial (''Tomistoma'') (and by inference other related extinct forms in Tomistominae) actually belong to Gavialoidea (and Gavialidae). Below is a clad ...
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Kentisuchus
''Kentisuchus'' is an extinct genus of gavialoid crocodylian, traditionally regarded as a member of the subfamily Tomistominae. Fossils have been found from England and France that date back to the early Eocene. The genus has also been recorded from Ukraine, but it unclear whether specimens from Ukraine are referable to ''Kentisuchus''. Species The genus ''Kentisuchus'' was erected by Charles Mook in 1955 for the species ''"Crocodylus" toliapicus'', described by Richard Owen, in 1849. William Buckland named ''"Crocodylus" spenceri'' on the basis of a partial skull found from the Isle of Sheppey in Kent, England. In 1888 Richard Lydekker considered ''"C." toliapicus'' synonymous with ''"C." champsoides'' and ''"C." arduini'', named by De Zigno, and reapplied the name ''"C." spenceri'' to all of these species. The genus name ''Kentisuchus'' was constructed only after it was realized that these specimens were clearly distinct from the genus ''Crocodylus'' and that some specimens ...
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Paratomistoma
''Paratomistoma'' (meaning "next to or near ''Tomistoma''") is an extinct monospecific genus of gavialoid crocodylian. It is based on the holotype specimen CGM 42188, a partial posterior skull and lower jaw discovered at Wadi Hitan, Egypt, in Middle Eocene-age rocks of the Gehannam Formation. The skull is unfused but considered morphologically mature. ''Paratomistoma'' was named in 2000 by Christopher Brochu and Philip Gingerich; the type species is ''P. courti'' in honor of Nicholas Court, who found CGM 42188. They performed a phylogenetic analysis and found ''Paratomistoma'' to be a derived member of Tomistominae, related to the false gharial. It may have been a marine or coastal crocodilian. Below is a cladogram based morphological studies comparing skeletal features that shows ''Paratomistoma'' as a member of Tomistominae: Based on morphological studies of extinct taxa, the tomistomines were long thought to be classified as crocodiles and not closely relat ...
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Toyotamaphimeia
''Toyotamaphimeia machikanensis'' (Toyotama-hime from Mountain Machikane ( :ja:待兼山)) is an extinct gavialid crocodylian which lived in Japan during the Pleistocene. A specimen recovered in 1964 at Osaka University during the construction of a new science building has been dated to around 430–380 thousand years old based on the stratum in which it was found. Unassigned species from same genus is also known from Taiwan. ''T. machikanensis'' was a fairly large crocodylian with a 1 m (3.3 ft) skull and a total length up to 7.7 m (25 ft). It was originally described as a member of the genus ''Tomistoma''. History and naming The first bones belonging to ''Toyotamaphimeia'' were discovered on May 3rd 1964 during the construction of a new school building on the grounds of Ôsaka University. A field survey was conducted shortly afterwards, confirming the presence of more fossils, however not yet identifying their crocodilian nature. Following the survey severa ...
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Gavialosuchus
''Gavialosuchus'' is an extinct genus of gavialid crocodylian from the early Miocene of Europe. Currently only one species is recognized, as a few other species of ''Gavialosuchus'' have since been reclassified to other genera. Taxonomy The type species, ''G. eggenburgensis'', is known from the early Miocene of Austria. Two other species - ''G. americanus'' and ''G. carolinensis'' - have since been reclassified to other genera. Myrick Jr. (2001) proposed synonymizing ''Gavialosuchus americanus'' with '' Thecachampsa antiqua''. Piras ''et al.'' (2007) advocated transferring both ''G. americanus'' and ''G. carolinensis'' to ''Thecachampsa'' as distinct species of the latter genus, however. Jouve ''et al.'' (2008) retained ''G. americanus'' in ''Gavialosuchus'' and found it to be the sister group of ''G. eggenburgensis'' (''G. carolinensis'' was not discussed). However, Jouve ''et al.'' (2008) didn't test ''Thecachampsa antiqua'' in their phylogenetic analysis. Shan ''et al.'' (200 ...
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Penghusuchus
''Penghusuchus'' is an extinct genus of gavialid crocodylian. It is known from a skeleton found in Upper Miocene rocks of Penghu Island, off Taiwan. It may be related to two other fossil Asian gavialids: ''Maomingosuchus petrolica'' of southeastern China and '' Toyotamaphimeia machikanensis'' of Japan. The taxon was described in 2009 by Shan and colleagues; the type species is ''P. pani''. Below is a cladogram based morphological studies comparing skeletal features that shows ''Penghusuchus'' as a member of Tomistominae, related to the false gharial: Based on morphological studies of extinct taxa, the tomistomines (including the living false gharial) were long thought to be classified as crocodiles and not closely related to gavialoids. However, recent molecular studies using DNA sequencing have consistently indicated that the false gharial (''Tomistoma'') (and by inference other related extinct forms in Tomistominae) actually belong to Gavialoidea (and Gavialidae). Below is ...
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False Gharial
The false gharial (''Tomistoma schlegelii''), also known by the names Malayan gharial, Sunda gharial and tomistoma is a freshwater crocodilian of the family Gavialidae native to Peninsular Malaysia, Borneo, Sumatra and Java. It is listed as Vulnerable on the IUCN Red List, as the global population is estimated at around 2,500 to 10,000 mature individuals. The specific name ''schlegelii'' honors Hermann Schlegel. Taxonomy The scientific name ''Crocodilus (Gavialis) schlegelii'' was proposed by Salomon Müller in 1838 who described a specimen collected in Borneo. In 1846, he proposed to use the name ''Tomistoma schlegelii'', if it needs to be placed in a distinct genus. The genus ''Tomistoma'' potentially also contains several extinct species like ''T. cairense'', ''T. lusitanicum'', ''T. taiwanicus'', and ''T. coppensi''. However, these species may need to be reclassified to different genera as are paraphyletic. The false gharial's snout broadens considerably towards the ...
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