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Maevarano Formation
The Maevarano Formation is a Late Cretaceous sedimentary rock formation found in the Mahajanga Province of northwestern Madagascar. It is most likely Maastrichtian in age, and records a seasonal, semiarid environment with rivers that had greatly varying discharges. Notable animal fossils recovered include the theropod dinosaur ''Majungasaurus'', the early bird ''Vorona'', the paravian ''Rahonavis'', the titanosaurian sauropod ''Rapetosaurus'', and the giant frog ''Beelzebufo''. Description The Maevarano Formation is well exposed in the Mahajanga Basin, in particular near the village of Berivotra near the northwestern coast of the island where its outcrops have been heavily dissected by erosion. At the time it was being deposited, its latitude was between 30°S and 25°S as Madagascar drifted northward after splitting from India about 88 million years ago. It is composed of three smaller units or members. The lowest is the Masorobe Member, which is usually reddish and is at least ...
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Geological Formation
A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics ( lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock exposed in a geographical region (the stratigraphic column). It is the fundamental unit of lithostratigraphy, the study of strata or rock layers. A formation must be large enough that it can be mapped at the surface or traced in the subsurface. Formations are otherwise not defined by the thickness of their rock strata, which can vary widely. They are usually, but not universally, tabular in form. They may consist of a single lithology (rock type), or of alternating beds of two or more lithologies, or even a heterogeneous mixture of lithologies, so long as this distinguishes them from adjacent bodies of rock. The concept of a geologic formation goes back to the beginnings of modern scientific geology. The term was used by Abraham Gottlob Wer ...
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Fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the ...
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Outcrop
An outcrop or rocky outcrop is a visible exposure of bedrock or ancient superficial deposits on the surface of the Earth. Features Outcrops do not cover the majority of the Earth's land surface because in most places the bedrock or superficial deposits are covered by soil and vegetation and cannot be seen or examined closely. However, in places where the overlying cover is removed through erosion or tectonic uplift, the rock may be exposed, or ''crop out''. Such exposure will happen most frequently in areas where erosion is rapid and exceeds the weathering rate such as on steep hillsides, mountain ridges and tops, river banks, and tectonically active areas. In Finland, glacial erosion during the last glacial maximum (ca. 11000 BC), followed by scouring by sea waves, followed by isostatic uplift has produced many smooth coastal and littoral outcrops. Bedrock and superficial deposits may also be exposed at the Earth's surface due to human excavations such as quarrying and build ...
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Beelzebufo
''Beelzebufo ampinga'' ( or ) was a particularly large species of prehistoric frog described in 2008. Common names assigned by the popular media include devil frog, devil toad, and the frog from hell. Fossils of ''Beelzebufo'' have been recovered from strata of the Maevarano Formation in Madagascar, dating to the late Cretaceous period, some 70 million years ago (Mya). It is considered to be closely related to ''Baurubatrachus'' from the Late Cretaceous of Brazil, with both possibly being close relatives, though not members of, the extant South American frog family Ceratophryidae. Discovery The first fossil bones were found in 1993 by David W. Krause of New York's Stony Brook University, but it took 14 years for scientists Susan E. Evans, Marc E. H. Jones, and Krause to assemble enough data for publication in the ''Proceedings of the National Academy of Sciences,'' the journal of the United States National Academy of Sciences. The generic name ''Beelzebufo'' is a portmanteau o ...
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Frog
A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" ''Triadobatrachus'' is known from the Early Triassic of Madagascar, but molecular clock, molecular clock dating suggests their split from other amphibians may extend further back to the Permian, 265 Myr, million years ago. Frogs are widely distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforest. Frogs account for around 88% of extant amphibian species. They are also one of the five most diverse vertebrate orders. Warty frog species tend to be called toads, but the distinction between frogs and toads is informal, not from Taxonomy (biology), taxonomy or evolutionary history. An adult frog has a stout body, protruding eyes, anteriorly-attached tongue, limb ...
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Rapetosaurus
''Rapetosaurus'' ( ) is a genus of titanosaurian sauropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period. Only one species, ''Rapetosaurus krausei'', has been identified. Like other sauropods, ''Rapetosaurus'' was a quadrupedal herbivore; it is calculated to have reached lengths of 15 metres (49 ft). Description ''Rapetosaurus'' was a fairly typical sauropod, with a short and slender tail, a very long neck and a huge, elephant-like body. Its head resembles the head of a diplodocid, with a long, narrow snout and nostrils on the top of its skull. It was a herbivore and its small, pencil-like teeth were good for ripping the leaves off trees but not for chewing. It was fairly modest in size, for a titanosaur. The juvenile specimen measured from head to tail, and "probably weighed about as much as an elephant". An adult would have been about twice as long ( in length) which is still less than half the length of its ...
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Sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their body), and four thick, pillar-like legs. They are notable for the enormous sizes attained by some species, and the group includes the largest animals to have ever lived on land. Well-known genera include ''Brachiosaurus'', ''Diplodocus'', ''Apatosaurus'' and ''Brontosaurus''. The oldest known unequivocal sauropod dinosaurs are known from the Early Jurassic. ''Isanosaurus'' and ''Antetonitrus'' were originally described as Triassic sauropods, but their age, and in the case of ''Antetonitrus'' also its sauropod status, were subsequently questioned. Sauropod-like sauropodomorph tracks from the Fleming Fjord Formation (Greenland) might, however, indicate the occurrence of the group in the Late Triassic. By the Late Jurassic (150 million yea ...
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Titanosaur
Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thriving at the time of the extinction event at the end of the Cretaceous. This group includes some of the largest land animals known to have ever existed, such as ''Patagotitan''—estimated at long with a weight of —and the comparably-sized ''Argentinosaurus'' and ''Puertasaurus'' from the same region. The group's name alludes to the mythological Titans of ancient Greek mythology, via the type genus (now considered a '' nomen dubium)'' ''Titanosaurus''. Together with the brachiosaurids and relatives, titanosaurs make up the larger sauropod clade Titanosauriformes. Titanosaurs have long been a poorly-known group, and the relationships between titanosaur species are still not well-understood. Description Titanosauria have the largest ...
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Rahonavis
''Rahonavis'' is a genus of bird-like theropods from the Late Cretaceous (Maastrichtian, about 70 mya) of what is now northwestern Madagascar. It is known from a partial skeleton ( UA 8656) found by Catherine Forster and colleagues in Maevarano Formation rocks at a quarry near Berivotra, Mahajanga Province.Tudge, Colin (2009) ''The Bird:A Natural History of Who Birds Are, Where They Came From, and How They Live'/ref> ''Rahonavis'' was a small predator, at about long and 0.45-2.27 kg (1-5 lbs),Holtz, Thomas R. Jr. (2008) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages'Supplementary Information/ref> with the typical ''Velociraptor''-like raised sickle claw on the second toe. It was originally the first African coelurosaur until the Nqwebasaurus was discovered in 2000. The name ''Rahonavis'' means, approximately, "cloud menace bird", from Malagasy ' (RA-hoo-na, "cloud" or "menace") + Latin ' "bird". The specific name, ''R. ostromi'', was co ...
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Paravian
Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the avialans, among which are the over ten thousand species of living birds. Early members of Paraves are well known for the possession of an enlarged claw on the second digit of the foot, which was held off the ground when walking in some species. Description Like other theropods, all paravians are bipedal, walking on their two hind legs. The teeth of primitive paravians were curved and serrated, but not blade-like except in some specialized species such as ''Dromaeosaurus albertensis''. The serrations on the front edge of dromaeosaurid and troodontid teeth were very small and fine, while the back edge had serrations which were very large and hooked. Most of the earliest paravian groups were carnivorous, though some smaller species (especially am ...
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Vorona
''Vorona'' ( ; Malagasy for "bird", ''V. berivotrensis'', "from Berivotra") is a monotypic genus of prehistoric birds. It was described from fossils found in a Maevarano Formation quarry near the village of Berivotra, Mahajanga Province, Madagascar. The age of the fossilised specimen is Late Cretaceous, probably Maastrichtian (72.1-66.0 mya). ''V. berivotrensis'' is known from scattered remains, possibly from a single individual ( UA 8651 and FMNH PA715). The phylogenic affinity of ''Vorona'' is hard to determine due to the fragmentary nature of the remains, mainly because the fossil shows a mix of basal avian features as well as some that seem very derived. ''Vorona'' might be a primitive ornithuromorph. At least two studies recovered it as part of Enantiornithes, however. ''Vorona'' is sometimes confused with the dromaeosaur ''Rahonavis ostromi'', a fossil of which was found in the same location. This confusion has led to the common misconception that ''Vorona'' had a dei ...
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Bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the bee hummingbird to the ostrich. There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming. B ...
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