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Macrocollum
''Macrocollum'' is a genus of unaysaurid sauropodomorph dinosaur that lived during the Late Triassic period (early Norian) in what is now Brazil. It is one of the oldest dinosaurs known. Discovery ''Macrocollum'' was discovered in 2012 in Rio Grande do Sul, Brazil, at the Wachholz site of the Candelária Formation, Paraná Basin. It was announced in a press conference on November 21, 2018. The generic name combines the Greek word μακρός (long) and the Latin word ''collum'' (neck), referring to the animal's elongated neck. The specific epithet honours José Jerundino Machado Itaqui, one of the main persons behind the creation of CAPPA/UFSM. Description Like most early dinosaurs, ''Macrocollum'' was relatively small, and walked on two legs. The known remains of ''Macrocollum'' are relatively well preserved. The holotype specimen consists of an almost complete and articulated skeleton. The two paratype specimens are both articulated skeletons with one missing a skull and i ...
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Macrocollum Pronto
''Macrocollum'' is a genus of unaysauridae, unaysaurid sauropodomorph dinosaur that lived during the Late Triassic period (geology), period (early Norian) in what is now Brazil. It is one of the oldest dinosaurs known. Discovery ''Macrocollum'' was discovered in 2012 in Rio Grande do Sul, Brazil, at the Wachholz site of the Candelária Formation, Paraná Basin. It was announced in a press conference on November 21, 2018. The generic name combines the Greek word μακρός (long) and the Latin word ''collum'' (neck), referring to the animal's elongated neck. The specific epithet honours José Jerundino Machado Itaqui, one of the main persons behind the creation of CAPPA/UFSM. Description Like most early dinosaurs, ''Macrocollum'' was relatively small, and walked on biped, two legs. The known remains of ''Macrocollum'' are relatively well preserved. The holotype specimen consists of an almost complete and articulated skeleton. The two paratype specimens are both articulated ske ...
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Candelária Formation, Paraná Basin
The Candelária Formation, in other literature also referred to as Candelária Sequence,Müller et al., 2017, p.543Martinelli et al., 2017, p.528 is a sedimentary formation of the Santa Maria Group (also called Santa Maria Supersequence) in the Paraná Basin in Rio Grande do Sul, southeastern Brazil.Candelária Formation
at .org
The formation dates to the of the

Unaysauridae
Unaysauridae is a clade of basal sauropodomorphs from the Late Triassic of India and Brazil. Diagnosis and systematics Unaysauridae was defined by Müller ''et al.'' (2018) as the most inclusive clade including ''Unaysaurus tolentinoi'', but not ''Plateosaurus engelhardti'' nor ''Saltasaurus loricatus''. Members of Unaysauridae are diagnosed by a substantially expanded cranial part of the medial condyle of the astragalus, as well as a promaxillary fenestra. Unaysauridae is sister to Plateosauria, more derived than ''Nambalia'', ''Thecodontosaurus'' ISI R277, ''Pantydraco'', and ''Efraasia''.Rodrigo Temp Müller; Max Cardoso Langer; Sérgio Dias-da-Silva (2018). "An exceptionally preserved association of complete dinosaur skeletons reveals the oldest long-necked sauropodomorphs". Biology Letters. 14 (11): 20180633. . ''Unaysaurus'' and ''Jaklapallisaurus'' had previously been assigned to Plateosauridae Plateosauridae is a family of plateosaurian sauropodomorphs from the Lat ...
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Sauropodomorph
Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The ''prosauropods,'' which preceded the sauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the Late Triassic (approximately 230 Ma) until their decline and extinction at the end of the Cretaceous. Description Sauropodomorphs were adapted to browsing higher than any other contemporary herbivore, giving them access to high tree foliage. This feeding strategy is supported by many of their defining characteristics, such as: a light, tiny skull on the end of a long neck (with ten or more elongated cervical vertebrae) a ...
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Unaysauridae
Unaysauridae is a clade of basal sauropodomorphs from the Late Triassic of India and Brazil. Diagnosis and systematics Unaysauridae was defined by Müller ''et al.'' (2018) as the most inclusive clade including ''Unaysaurus tolentinoi'', but not ''Plateosaurus engelhardti'' nor ''Saltasaurus loricatus''. Members of Unaysauridae are diagnosed by a substantially expanded cranial part of the medial condyle of the astragalus, as well as a promaxillary fenestra. Unaysauridae is sister to Plateosauria, more derived than ''Nambalia'', ''Thecodontosaurus'' ISI R277, ''Pantydraco'', and ''Efraasia''.Rodrigo Temp Müller; Max Cardoso Langer; Sérgio Dias-da-Silva (2018). "An exceptionally preserved association of complete dinosaur skeletons reveals the oldest long-necked sauropodomorphs". Biology Letters. 14 (11): 20180633. . ''Unaysaurus'' and ''Jaklapallisaurus'' had previously been assigned to Plateosauridae Plateosauridae is a family of plateosaurian sauropodomorphs from the Lat ...
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Jaklapallisaurus
''Jaklapallisaurus'' is a genus of unaysaurid sauropodomorph dinosaur. It lived during the Late Triassic period (late Norian to earliest Rhaetian) in what is now Telangana, central India. It is known from the holotype ISI R274, postcranial material which was collected from the Upper Maleri Formation (late Norian–earliest Rhaetian) of the Pranhita–Godavari Basin and from the referred material ISI R279, partially complete right femur which was collected from the Lower Dharmaram Formation (latest Norian–Rhaetian). It was first named by Fernando E. Novas, Martin D. Ezcurra, Sankar Chatterjee and Tharavat S. Kutty in 2011 and the type species is ''Jaklapallisaurus asymmetrica''. The generic name is derived from the Indian town of ''Jaklapalli'' which is close to the type locality. The specific name refers to the highly asymmetrical astragalus of this species in distal view. A cladistic analysis by Novas et al. found that all valid plateosaurid species form a large p ...
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Unaysaurus
''Unaysaurus'' is a genus of unaysaurid sauropodomorph herbivore dinosaur. Discovered in southern Brazil, in the geopark of Paleorrota, in 1998, and announced in a press conference on Thursday, December 3, 2004, it is one of the oldest dinosaurs known. It is closely related to plateosaurid dinosaurs found in Germany, which indicates that it was relatively easy for species to spread across the giant landmass of the time, the supercontinent of Pangaea. The fossils of ''Unaysaurus'' are well preserved. They consist of an almost complete skull, complete with a lower jaw, and partial skeleton with many of the bones still connected to each other in their natural positions. It is one of the most complete dinosaur skeletons (including complete skull) ever recovered in Brazil. Discovery and naming ''Unaysaurus'' was found in the southern Brazilian state of Rio Grande do Sul, near the city of Santa Maria. It was recovered from the red beds of the Caturrita Formation, which is the geo ...
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Million Years Ago
The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). Together they make a reference system, one to a quantity, the other to a particular place in a year numbering system that is ''time before the present''. Myr is deprecated in geology, but in astronomy ''Myr'' is standard. Where "myr" ''is'' seen in geology it is usually "Myr" (a unit of mega-years). In astronomy it is usually "Myr" (Million years). Debate In geology a debate remains open concerning the use of ''Myr'' (duration) plus ''Ma'' (million years ago) versus using only the term ''Ma''. In either case the term '' Ma'' is used in geology literature conforming to ISO 31-1 (now ISO 80000-3) and NIST 811 recommended practices. Traditional style geology literature is written The "ago" is implied, so that any such year number "X Ma" ...
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Faunal Stage
In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by convention have the same name, and the same boundaries. Rock series are divided into stages, just as geological epochs are divided into ages. Stages can be divided into smaller stratigraphic units called chronozones. (See chart at right for full terminology hierarchy.) Stages may also be divided into substages or indeed grouped as superstages. The term faunal stage is sometimes used, referring to the fact that the same fauna (animals) are found throughout the layer (by definition). Definition Stages are primarily defined by a consistent set of fossils (biostratigraphy) or a consistent magnetic polarity (see paleomagnetism) in the rock. Usually one or more index fossils that are common, found worldwide, easily recognized, and limited to a sing ...
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Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', t ...
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Triassic
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic. The Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Three categories of organisms can be distinguished in the Triassic record: survivors from the extinction event, new groups that flourished briefly, and other new groups that went on to dominate the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized subgroup of archo ...
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Suture (joint)
In anatomy, fibrous joints are joints connected by fibrous tissue, consisting mainly of collagen. These are fixed joints where bones are united by a layer of white fibrous tissue of varying thickness. In the skull the joints between the bones are called sutures. Such immovable joints are also referred to as synarthroses. Types Most fibrous joints are also called "fixed" or "immovable". These joints have no joint cavity and are connected via fibrous connective tissue. The skull bones are connected by fibrous joints called '' sutures''. In fetal skulls the sutures are wide to allow slight movement during birth. They later become rigid ( synarthrodial). Some of the long bones in the body such as the radius and ulna in the forearm are joined by a ''syndesmosis'' (along the interosseous membrane). Syndemoses are slightly moveable ( amphiarthrodial). The distal tibiofibular joint is another example. A ''gomphosis'' is a joint between the root of a tooth and the socket in the max ...
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